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Regarding taxon authors in lists. I would be tempoted to propose a fairly strict policy, because this mess is getting out of hand. I am trying to get Catocala a better "Well-written" rating and naturally one would improve list layout to make it more easily legible. Now, it is an absolute pain to look for a taxon.
Doing so, I noted an inconsistency rate of taxon author/dates between list and taxoboxes of about 15%. Didn't count it, but I'd safely say it's between 20 and 10%. That is too fricking high for Wikipedia. If would mean that every 10th word at least in an article is slightly but (for the average user) invisibly and insiduously wrong. Now, even Encyclopedia Britannica achieves higher standards. By Wikipedia standards, it is abysmal. The list, the taxonboxes, they simply cannot be trusted.
The present proliferation of listing taxon author/date is apparently not manageable and must probably be curtailed. Severely. (It used to be more restrained, and we did not have so many errors back then)
Rationale: What use is author/date? Usually not much, considering the price we pay for having it everywhere:
In scientific writing, author/date are usually left out; this is the convenient way and should be done so in casual writing also. However, they are usually cited at least once somewhere in an article. Proper style is to cite them or discuss them where important or most important. In practice, this is either within a taxonomic narrative, a taxonomic list, or in case of ambiguities such as homonyms, synonymy etc. But zoology papers that are not about nomenclature or taxonomy often do not give them anywhere (even "pure" systematics/phylogenetics papers sometimes do). In botany, giving the authorship at least once somewhere is essentially mandatory due to the homonymy proliferation; even some botany authors leave out the data entirely however. And when scientists write author/date outside a taxon list or table -- i.e., if they are inserted into a running paragraph -- it is considered good style to smallcaps them or similar.
The general public has a hard time citing taxa correctly ("T. Rex" syndrome). Citing author/date in a casual context outside a specific (scientific) setting is stylistic hypercorrection; popular writers do it more often than scientists. But it is not professional. Wikipedia should not endorse bad writing style.
Generally, except in case of homonymies, author/date of taxa are fairly useless to the standard user. Interesting trivia ("how many [taxon] were described after 2000?"), but Wikipedia is explicitly not a collection of what is equivalent to statistical trivia. We should to have the data somewhere of course, but we do not to have it more than once, especially if that is at the mot obvious location (i.e. the taxobox).
The present author/name inconsistencies must be sorted out sooner or later anyway. The quickest way to do so is to collect all such information at one point. At present however, nomenclatorial data on WP:TOL is in need of complete review.
While most errors are small (parentheses wrong, date not correctly determined, authors ambiguous) and easily resolved, perhaps 10% are serious. Keeping it all accurate would require constant maintenance effort, and at some point the workload exceeds the workload we can put up. (Arguably, this is already the case.)
It wastes space. A lot of it. And we need every bit of space if we want to have as little as comprehensive "scientific name + common name" lists for diverse taxa. Genera with 150 species, it's everyday occurrence in beetles and moths.
It makes reading the text harder; it is a useless baroque flourish in most contexts, especially if given in the very first sentences.
Sources are generally pieces of shit. Seriously. A list one finds on the internet is not a taxonomic/nomenclatorial source; it is not even a WP:RS. There are the good web databases/lists everyone knows for their taxon, museum databases etc, but these are often enough faulty also. There is one and only one completely reliable source for taxon author/date: the original description.
In plain English:
Wikipedia has become the largest, and worst, source in the world for nomenclatorial data. "Worst" referring to both accuracy and style-of-presentation.
Hence, I ponder proposing something along the lines of:
"No taxon author/date is given, except 1. at redlinks, 2. in taxoboxes, and 3. where it is pretty damn important (homonymies, mostly)." (This implies that all author/date info we have is moved up the tree towards the pertient taxoboxes as far as possible. E.g. if we only have the family article with a redlist of genera, the genera get author/date. When the genus articles are made, author/date move to the genus article taxobox, and the process is repeated for species (red)links)
"Taxon author/date refs MUST be the original description. Stuff "someone found on the Web" goes in "External links"." (Yes, even "known good lists" - even IUCN, Fauna Europaea, WoRMS, biolib.cz, funet.fi, GRIN etc have 5-10% error rate. Claiming a source but actually having only a crap source belies the principle that makes Wikipedia work: Readers trust our sources to be good WP:RS. They do not habitually read the fulltext source, they usually just check out that "sources exist" in case they ever need one. Additional rationale: if we do this, we do have the orig. descr., for whatever we want to do with it. Which can be quite a lot, and for many spp. is actually all we will ever get.)
(Additional guideline: If taxon author/date are to be mentioned in the article maintext, do it as a proper narrative. E.g. at the end of the intro section, or in the taxonomy Systematics and nomenclature/Classification section something like "Blah blub was first scientifically described (as Bulb blub) by Mork Schmorkus in 1796, and put in its present genus by Hank Kolf in 1999."
If you really prefer to simply throw a "The Great GumbletreeBlah blub Schmorkus (Kolf), 1796 (1999) is a tropical rainforestcanopy treeendemic to Belize" at the unsuspecting reader, at least make it ((aut|Schmorkus (Kolf), 1796 (1999))) or <small>Schmorkus (Kolf)</small> or something similar. Please.
PS: Remember the rules of biological nomenclature do not require you to give author/date within a narrative. It will be in the taxobox anyway, that's what the taxobox is there for (among other things). Remember also that just beacuse you habitually read over stuff like "The Great Gumbletree Blah blub Schmorkus (Kolf), 1796 (1999) is a tropical rainforest canopy tree endemic to Belize", most people out there would more likely think "WTF?!" instead of understanding much of it. Most do not even have English as a first language, let alone Scientiese.) Dysmorodrepanis (talk) 21:53, 28 March 2012 (UTC)
I generally don't put the author (again note that most botanical disciplines with a few exceptions don't include the date) anywhere except on lists of taxa and in the taxobox. I'll have to think more about your point 1, but on point 2, I completely disagree. Original descriptions are often crap, especially if they're from the early 1800s. I was looking at one the other day and it's a simple lineup of new names with a very short Latin description, no author information except that implied by the name on the book. And sometimes the author is attributed incorrectly for a couple centuries. Only recent analysis of supporting manuscripts and letters allowed Juliet Wege to determine that the authority of Stylidium is not Sw. ex Willd., but just Sw. (see [1]). Further, not everyone has access to the original publication, so it shouldn't be cited when the editor hasn't seen the source: WP:SAYWHEREYOUGOTIT. I agree that online sources like ZipCodeZoo and the Plant List and even EOL are often terrible sources for authors. IPNI even has its faults, but it's mostly correct, which is why I cite it frequently. IMHO, I think we just need to encourage editors to get this information right from reliable sources, principles we already have and are well supported. This doesn't seem to be a specific problem to author data, but a general plea to use the best reliable sources you can find. Rkitko (talk) 00:38, 29 March 2012 (UTC)
Citing the original description may be useful sometimes, but it is certainly not enough. When we quote the authority, we are not just stating that that person described such a species, but that theirs is the accepted, valid name, which normally, but not always, means it was the first description of the species. It may have been later rejected, it may be a junior homonym, or any number of other exceptions may apply. That's why we need a third-party source such as WoRMS, IUCN, etc., to confirm that the authority is correct. I generally only cite the original description for details of the morphology or ecology, not for the authority itself. I don't see that your first point is really fixing a serious problem. Small caps are explicitly forbidden from such cases, and I've had to spend a lot of time in the past removing ((aut)) that you (Dysmorodrepanis) have added; you're pretty much the only one, and it would be preferable if you could stop. It's definitely not something to be promoted. --Stemonitis (talk) 01:45, 29 March 2012 (UTC)
I would agree that authorities are not best placed in running text, at least on Wikipedia, but I would also second everything that Rkitko said; if we use the best sources, accuracy increases. --Stemonitis (talk) 01:59, 29 March 2012 (UTC)
We do NOT in fact use the best sources; the rule you propose is proven unenforceable! The third-party source is the appropriate Code of Nomenclature, which sets unequivocal rules that no sloppy database administrator or weblist compiler is allowed to violate. And Wikipedia editors are not allowed to volate them either. Dysmorodrepanis (talk) 14:37, 29 March 2012 (UTC)
I can't follow this argument at all. You think that instead of citing, say, WoRMS, we should cite the ICZN, which almost certainly doesn't mention the name in question? Even if a name is invalid according to the code, if it is used by reliable sources, then we must follow it. For instance, if one of us discovers a case of homonymy, then we have to continue to use the invalid homonym until a replacement is published elsewhere. It can be one of us who publishes it (not unheard of), but it cannot be introduced on Wikipedia. The codes limit us only indirectly in that they limit what appears in reliable sources. Avoiding original research is a general principle across Wikipedia, and I'm surprised that it's being misunderstood here. --Stemonitis (talk) 14:55, 29 March 2012 (UTC)
As others have noted, there is a major fallacy in the use of the original publication as the source for the authority of a taxon: how do you know which is the original publication? The only way you can decide on this is to use a secondary reliable source which uses taxonomic research to discover the answer to this question. Hence this secondary RS is the actual source. If I write that the authority for the name X y is Jones (with or without a date), I need to cite a source that says that Jones was indeed the authority for this name; the publication by Jones is useless for this purpose – it doesn't tell you whether the name was properly published according to the relevant code. At best the original publication tells you that Jones claimed to have established the name X y, not whether Jones succeeded. This statement "There is one and only one completely reliable source for taxon author/date: the original description" is simply wrong.
(In fact, using the original publication only is a violation of WP:PRIMARY: "All interpretive claims, analyses, or synthetic claims about primary sources must be referenced to a secondary source". By using the original publication as the only source, you're making a judgement that it succeeded according to the relevant code and you need a source for this judgement.)
Wikipedia is as much bound by the Codes of Nomenclature as anyone else except Creationists and other intellectual lowlife. What constitutes the source for a taxon is not determined by a secondary source or anyone's interpretation, but by mandatory rules (Chapters 3-4 and 11 of the ICZN Code, Chapter IV of the ICBN) to which everyone IS REQUIRED to stick. And these rules do not grant any space to "interpretations"; they were designed to prtevent exactly what you fear. Thus, WP:PRIMARY is not violated. In fact, it may be argued that we must use the original description only, because WP:RS overrules everything else regarding sources, and there is only one truly reliable source for taxon author/dates: the original description. The "'no' primary sources" rule serves to prevent ambiguous and subjective interpretations, whereas the ambiguity in the present case is caused by the secondary (and especially tertiary) sources! Here, the primary source is always right (except in gross spelling errors, if these are corrected later - in which case we do have the secondary source at hand already). And in any case, as soon as a conflict between secondary/tertiary sources occurs (and there are literally thousands of them, those databases are notoriously sloppy when it comes to disambiguating authors such as "Meyer"s or distinguishing date-of-issue from date-of-valid-publication (anything published in the PZSL is liable to be mis-dated), we have to turn to the original description anyway! E.g. for determining whether date-of-publication requires square brackets or not, the primary source MUST be considered (and cited).
Consider the case of legal documents, such as the Uniform Code of Military Justice. Here, the primary source is used throughout. Since the Codes of Nomenclature are equivalent to "law", this is a close parallel. As an example closer to biology, consider Copernicium#Naming, where the pertinent IUPAC documents (primary sources) are cited with nobody batting an eye. Your reservations against primary sources make no sense in any field where stuff is decided by mandatory rules; hence the exceptions in WP:PRIMARY. (For legal issues, we are even forced to add a disclaimer to cover the problems caused by secondary and tertiary sources, because only the primary wording of laws is binding and authoritative, not their subsequent interpretation!) (talk) 14:37, 29 March 2012 (UTC)
I'm sorry, but your analogy is quite wrong. If the ICZN and ICBN listed all accepted names within them then of course they would be the authoritative sources. But they do not. They contain the rules which apply to names, not the names themselves (other than a relatively small proportion in examples or conserved names, etc.). The ICZN and ICBN are authoritative as regards the rules (and any names they specifically list). The application of the rules in deciding whether all other names are ok needs to be sourced. The precise analogy seems to be using a legal document to decide whether or not a particular trial or judicial decision was correct. This would be wrong. Peter coxhead (talk) 15:46, 29 March 2012 (UTC)
I was nowhere talking about taxon names. I was talking about author names and authorship dates. E.g. which "[Schiffermüller], 1775", "Schiffermüller in Denis & Schiffermüller, 1775" or "Denis & Schiffermüller, [1775]" -- if any-- is correct. What to do if one source gives "(L.) Blume" and another "(L.f.) Blume" and a third "L.". Dysmorodrepanis (talk) 22:13, 29 March 2012 (UTC)
I completely agree that authorities should not be included in running text as a matter of course, but only in taxoboxes and lists. On the other hand, in some areas of taxonomy, there is so much muddle that only by including authorities can you disambiguate. To take just two I've worked on lately: Epiphyllum Haw. is a different genus to Epiphyllum Pfeiff.; Ophrys holoserica auct. is a different species to Ophrys holoserica (Burm.f) ex Greuter. Of course there are ways of saying this other than writing the Latin name followed by the authority, and these are clearly better for the general reader, but the authority sometimes needs to be mentioned, even in running text.
This is precisely one of the 3 exceptions I mentioned in the suggestion (the third one, "pretty damn important"). Dysmorodrepanis (talk) 14:37, 29 March 2012 (UTC)
Small caps There is continuing discussion of deleting the ((smallcaps)) template to which ((aut)) is a redirect. MOS:SMALLCAPS is quite clear: "Change small caps to title case." ((Aut)) is a monstrosity and should not be used. Peter coxhead (talk) 08:49, 29 March 2012 (UTC)
In proper scientific writing, this "monstrosity" is used regularly. If Wikipedia is so stupid as to reject what everyone else in the business accepts, then to hell with its respectability among the scientific community, which a lot of people have worked damn hard to build in the first place! Matter of fact, I cannot find anything at Template talk:Smallcaps to support your claims. Dysmorodrepanis (talk) 14:37, 29 March 2012 (UTC)
Try looking at Template:Smallcaps: "small caps should be avoided and reduced to one of the other title cases or normal case and markup should be kept simple". --Stemonitis (talk) 14:55, 29 March 2012 (UTC)
I tried looking at Wikipedia:Templates_for_deletion/Log/2008_December_2#Template:Smallcaps instead. The result of the discussion is unequivocal. In any case, I would not recommend to use smallcaps here; I'd rather completely avoid citing author/date in a way where professional authors would use smallcaps. Make it a narrative instead, as per example given. Dysmorodrepanis (talk) 15:06, 29 March 2012 (UTC)
Some professional authors would use small caps, perhaps, although I have to say that I can't recall seeing it in the botanical literature. Peter coxhead (talk) 15:46, 29 March 2012 (UTC)
And I recall seeing it a lot in the zoological literature. It probably depends on the journal. Dysmorodrepanis (talk) 22:08, 29 March 2012 (UTC)
It may be helpful to consider the rules by which we are bound:
51.1. Optional use of names of authors. The name of the author does not form part of the name of a taxon and its citation is optional, although customary and often advisable.
Recommendation 51A. Citation of author and date. The original author and date of a name should be cited at least once in each work dealing with the taxon denoted by that name. This is especially important in distinguishing between homonyms and in identifying species-group names which are not in their original combinations. If the surname and forename(s) of an author are liable to be confused, these should be distinguished as in scientific bibliographies.
46.1. In publications, particularly those dealing with taxonomy and nomenclature, it may be desirable, even when no bibliographic reference to the protologue is made, to cite the author(s) of the name concerned [...]
In a nutshell, they suggest a "use it sparingly, but use it at least once except in case of homonymies etc. where it should be used whenever there is risk of confusion" policy.
As a case in point, Noctua (moth). This is what the current SOP will get us: 10 taxa with articles, 50% of these have an author/date conflict between genus and species article, and all that have common names have the wrong citation style.
Do you want this to proliferate? Anyone who says "yes" is herewith kindly asked to go ahead and fix up this mess. Dysmorodrepanis (talk) 14:37, 29 March 2012 (UTC)
Reply to point 8: There are sources and sources. One must make a distinction between sources made by professional zoologists or botanists and other sources. As stated, even trusted sources sometimes contain mistakes. Then it is up to us, when we find a mistake, to send an email to that source explaining the problem. I have done this many times and have always received a polite response. The problem then was soon taken care of. JoJan (talk) 14:46, 29 March 2012 (UTC)
The scope of the problem prevents this approach. Do you want to check all 447449 taxa currently on WoRMS? (The fact that WoRMS has a very low error rate makes it worse; it is simply a waste of time.) Dysmorodrepanis (talk) 15:00, 29 March 2012 (UTC)
I agree with Stemonitis somewhere above in this mess; it shouldn't be Wikipedia editors' jobs to interpret the ICZN (which is frequently open to different interpretations anyway: what is "prevailing usage"?). The difference with Copernicium is that in that case the IUPAC dealt with this specific name; the ICZN deals only with generalities that have to be applied to specific cases. We should follow secondary sources, such as major taxonomic reviews and well-maintained databases like WoRMS. Ucucha (talk) 15:05, 29 March 2012 (UTC)
Again: the secondary sources regularly (though inadvertently) violate the Codes of Nomenclature by which we are bound. Hence, they cannot generally qualify as WP:RS; their reliability must theoretically be proven again and again in each and every case. The Codes are also not subject to interpretation with regard to the points discussed here. In fact, they could hardly be more unequivocal. Where exactly is space for subjective interpretation in a statement such as:
The valid name of a taxon is the oldest available name applied to [its type specimen(s)], unless that name has been invalidated or another name is given precedence by any provision of the Code or by any ruling of the Commission.
That is what we are considering here, and it is laid out cleanly and unequivocally. Whether some taxon is considered valid - now THAT is something where we have to turn to secondary sources (if such are available). But we do that already, and nobody proposes changing it. The question here is rather: 1. who first validly gave that name to that taxon and 2. when did that happen? Here, only the primary source is certainly a WP:RS (printer's errors nonwithstanding, but for these we will turn to secondary souces as usual). The problem is that the dating is done somewhat contrary to everyday practice (Art. 21 ICZN Code), and many database or Web list compilers do not know this or do not take enough care (the sqare-brackets problem, perhaps the most common nomenclatorial error on Wikipedia) Dysmorodrepanis (talk) 15:32, 29 March 2012 (UTC)
@ Dysmorodrepanis: I check the taxa as I go along. And I only check what is inside my scope of expertise, not the whole database. If we all do the same, then, in the long run, we have a win-win situation. Wikipedia has correct information and the database stands corrected. JoJan (talk) 15:14, 29 March 2012 (UTC)
I do the same, but the problem is: no matter how much time we put into maintenance/quality control, it is not sufficient. Back in 2007/2008, you could blindly trust the author/name data on Wikipedia; we hade not that many articles then, and those we had were usually well-known taxa. Today, there are so many articles on little-known taxa which contain so many hard-to-check errors that Wikipedia is close to useless as a nomenclatorial point-of-info. Even if some external source is given, you cannot tell whether it's correct. We are fighting a losing battle here; consequently the system must change, not the way we deal with it. Dysmorodrepanis (talk) 15:32, 29 March 2012 (UTC)
(NB: taxonomic databases are not usually secondary nomenclatorial sources. To compile a secondary source, the primary source must be considered directly. Database authors usually do not do this. Rather, they compile the information from check-lists or similar, and they usually do not do error checking either. Consequently, the databases are tertiary sources (or even quaternary sources, if such a distinction is made), and this kind of throws a lot of what is said above out of the window... (we are advised to use tertiary sources with as much restraint as primary sources). The default secondary source in nomenclature is the "first reviser" work, ICZN Code Article 24.2.). Dysmorodrepanis (talk) 15:46, 29 March 2012 (UTC)
Where exactly is space for subjective interpretation in a statement such as:
The valid name of a taxon is the oldest available name applied to [its type specimen(s)], unless that name has been invalidated or another name is given precedence by any provision of the Code or by any ruling of the Commission.
"Subjective" interpretation isn't the issue. How do you know that a name is the "oldest"? How do you know that it was "available"? This can only be determined by original research in the taxonomic literature. I can only repeat my previous example. You find an original publication by Jones in 1800 which names species X y. This does not tell you that this was the oldest; this does not tell you that the name was available; this does not tell you that the naming was validly done under the relevant code.
Our task is to use the most reliable secondary sources that we can. We can certainly check with primary sources for straightforward mistakes (e.g. I've found transpositions in dates such as "1953" for "1935"), but we shouldn't ourselves be making judgements based on the taxonomic codes (although WP:IAR applies in exceptional cases). Peter coxhead (talk) 16:02, 29 March 2012 (UTC)
You talk about taxon names, and basically everything you say is right. But it completely misses the point, I was talking about author names and description dates. By having an article at all, we need (technically) a secondary souce. This decides all that taxonomic stuff you mention. How the correct way of writing author/date is determined is decided by the ICxN.
(There will be the occasional case of nomenclatorial dispute, the type specimen of Archeopteryx)
Again: taxonomy databases are usually not secondary sources for nomenclatorial questions; They are usually not based on primary sources. Hence, they are tertiary sources, which we cannot use freely either.
As a final comment, the underlying argument Dysmorodrepanis uses is odd: because there are errors in authorities/dates, articles shouldn't normally contain any or should at least drastically reduce the number they contain. I'm rather more concerned about the many more obvious and substantive errors in biology-related articles; so should I propose that we urge editors to stop producing such articles? After all, there are so many that we'll never be able to fix them all... Peter coxhead (talk) 16:08, 29 March 2012 (UTC)
It is not odd, nomenclatorial factoids are not the same as article "bread and butter"; on the contrary, "statistical" minutiae are explicitly forbidden as wikipedia content (Wikipedia:NOTSTATSBOOK). We need to have the information, it is an integral part of the taxon "as a dataset". But we do not need to have it where it increases the total number of errors while adding little other than bulk; Wikipedia standards argue against it and the ICxN does not argue for it either except in the 3 exceptions I drew up above.
Also, it may be worthwhile to leave the colorful bits to new editors, there they can learn their mettle. Us veterans are probably better off doing grunt work, unfortunately - Wikipedia editor growth is slowing, and one reason is that the environment is not as attractive anymore as it was.
Two things got mixed up a bit from what I originally intended:
The issue at hand (bad data), the "Rationale" above, and the 2 points below it are all different things. The main thing is: I am not the only one who recognizes a problem - how do we solve it?
Read #Taxonomy_vs_Classification_vs_Systematics_vs..... above if you haven't done so. The problem is not about secondary sources vs primary sources. Whether a taxon is valid taxonomically (whether we can have an article or not), that is and always will only be up to the secondary sources to decide. If these say "it is valid", we must decide on authorship data, how exactly it is written correctly, which is purely nomenclatorial data, to which external "laws" apply which are quite unequivocal. Here we have primary sources alyways, secondary sources rarely, tertiary sources always.
Here is the problem: a proliferation of doubtfully reliable tertiary sources -- weblists and databases --, when 99.9999999% reliable primary sources exist, but most secondary sources are in fact long monographs and lists printed before 1970. Catalogue of the Birds of the British Museum, that would be an example of a secondary nomenclatorial source. Remember the long "old-fashioned" synonymy lists they have there? Typical for secondary nomenclatorial sources. These were compiled from the primary sources. What the databases and weblists have has been copied half a dozen times or more. Every time, a lapsus may occur.
I have observed that if you take all the "good" tertiary sources now in use for nomenclature on Wikipedia together, you have for perhaps up to 50% (certainly not less than 20%, I'd say) of the taxa more than one version of the nomenclatorial data. One or none of these are correct. All others are lapsus. If WoRMS says one thing and Fauna Europaea another, and IUCN sides with the second, but some museum catalogue sides with the first, and ITIS introduces yet another variant, who are we gonna believe? What if GRIN and ILDIS contradict each other? Go with ILDIS, OK, but still 25% chance you're wrong. It may only be obvious who copied whom, but it will never be obvious who is correct unless we consult the primary source. Citing one tertiary source simply isn't enough often enough. We'd need to check out at least 3; if they agree we're likely in the clear. Some (mainly plant) articles already do so (with plants, I'd guess it's out of necessity.); it slows down browsing and wastes space.
My solution presently is roughly a) collecting/collating most (except homonyms etc) of the data at the smallest number of specific points -- the most specific taxobox or intext list by cutting and then pasting any differences), and/or b) generally eschewing tertiary sources for the primary source (secondary in case of amendations to the original nomens) for taxon authorship data -- since in any case of dispute we need to turn to the primary data anyway, we may just as well use it as the first source to check out, as well as b1) generally having no citations for author/date
Anyone perchance got time to read through Stemmatics? Documentary phlogenetics, getting a few ideas from there might help us.
What I would really like to propose:
Considering we basically agree that there might be a problem requiring fixing, we should do an improv study. Science can be "hard" or "soft", and there is also pseudoscience ;-)
Hard: Take random selection of perhaps 200 or more taxon articles (i.e. taxobox template present). Take their parent taxon articles. Take the parent taxon articles of those. If the articles selected are not highest-rank (species/subspecies), do the same one step downwards. Compare authorship data, note any discrepancies. Half a year later, do the survey again, for a) the original sample of articles and b) a new random sample. Compare results.
Soft: Routinely do such comparisons one taxonomic level up/down, whenever you do major edits on taxon articles. Note down how many % discrepancies. Half a year later, compare collected data.
Pseudo: Take two random 18th-century nomenclatorial works, e.g. Systema 12th ed. Take all species described therein which are on Wikipedia. In the species taxoboxes and on any genus lists where authorship data is given, count parentheses errors -- for the Systema, "Linnaeus, 1766" where it should be "(Linnaeus, 1766)" (the reverse probably does not occur) or "L." where it should be "(L.) [reviser]". Also, count date errors -- "1766" where it should be "1767" (the reverse probably does not occur either). Wait half a year, repeat, compare.
OK. Thanks for your attention.
Now, any volunteers to fix Cochranella? It very much illustrates all the points I have brought up.
We apparently have 3 databases in there now: IUCN (the species articles), another one at the initial version of the genus article -- perhaps ITIS (I moved the data from genus to species long ago, hence the "verification needed"s at many species already), plus, apparently, a third (someone restored author/dates at the genus list, but some "verification needed" species do not disagree with the new list. Dysmorodrepanis (talk) 21:46, 29 March 2012 (UTC)
Biota vs species
There has been a discussion on using the term "biota" as opposed to "species". Has this been resolved? Category:Endangered species has a mish-mash of "endangered species" and "endangered biota". -- Alan Liefting (talk - contribs) 23:32, 29 March 2012 (UTC)
Where have this been discussed? Petter Bøckman (talk) 06:37, 30 March 2012 (UTC)
In this particular instance (i.e. the context of "endangered") the issue is that a small proportion of the taxa listed as endangered are subspecies, not species, so "endangered species" isn't right as a category name. Why "endangered biota" is better than "endangered taxa" isn't clear to me. It would certainly be better to be consistent throughout the categories of endangered groups. Peter coxhead (talk) 08:31, 30 March 2012 (UTC)
Got the point. I would use "groups" or "taxa", particularly since the term "Biota" already has an established meaning in biology. Petter Bøckman (talk) 11:04, 30 March 2012 (UTC)
"Endangered species" is the commonly used term. On the principle of least surprise it should be used instead of "endangered biota", even though the latter may be a more accurate description. -- Alan Liefting (talk - contribs) 22:17, 30 March 2012 (UTC)
I suppose "endangered biota" is the proper term when a whole eco-system is under threat. At least in ecology, biota is the sum total of all organisms in the system under discussion. I have never heard it used for single species. Petter Bøckman (talk) 22:24, 30 March 2012 (UTC)
I think we can all agree that "endangered biota" isn't right. I understand Alan's point but on the other hand he has rightly argued for precision and accuracy in categorization and I'd be unhappy to have to categorize an endangered subspecies as [[Category:Endangered species]]. Because laypeople use "species" incorrectly and therefore wouldn't be surprised isn't a wholly convincing reason for us to do so, at least in my view. Peter coxhead (talk) 22:41, 30 March 2012 (UTC)
A further note on sources
1. Why taxonomic databases are (mostly) tertiary sources for nomenclature, but may be legitimate secondary sources for taxonomy:
As per Markku Savela (emphasis changed):
Taxonomic referencing Don't do it! Or, if you do, keep in mind that this site is a collection of names (taxa). The status or ranking of a specific taxon in this site should not be referenced. This site does not contain orignal taxonomic information or opinions. Everthing is based on published literature or other information given to me.
But many of us (including myself) use this site as a mainstay ref. For WoRMS, basically the same disclaimer (in Savela's original emphasis, see link) holds true -- its taxonomic structure as is is highly original research (emphasis added):
As a user or developer you can use the WoRMS webservice to feed your own application with standard WoRMS taxonomy.
The same holds true for Fauna Europaea and ABRS: editors have considerable taxonomic leeway leafwards in the tree.
Theritas and others
Now, consider for example Theritas. Second section from bottom is the list of main references. You will note that these range from primary/secondary (BCA, Hemming etc) to "beyond tertiary" (Smart). FE, ABRS, WoRMS etc use more and less tertiary sources, but it is obvious that they are mostly built on previously-published (secondary) lists, not on doing species by species from the primary literature.
2. Primary vs tertiary sources
We should use both with restraint. While this is remembered for primary sources (or at least it is claimed, the reality is different), it is less openly remembered for tertiary sources, which are defined as
publications such as encyclopedias and other compendia that summarize primary and secondary sources.
This includes most taxonomic databases. Compare:
Everthing is based on published literature or other information given to me.
There is, however, WP:SCHOLARSHIP. A peer-reviewed publication in a journal that is not disreputable (no exact definition, but "you know it when you see it") is for WP purposes considered RS until proven otherwise. While 1' and 3' sources are acceptable under some circumstances (WP:NOR etc), they better be scholarly + peer-reviewed. Luckily, most nomenclatorial acts fall into this category, as do most taxonomic acts. The former are by definition 1' sources, the others may be anything but are mostly 3' sources these days (a taxon is proposed 1', it is vetted 2', it is accepted or rejected 3').
My remarks are not concerned with using 3' sources for taxonomy. This is fine enough (though I prefer secondary peer-revewed). It is about the SOP using 3' sources to determine the correct author/date citation of the nomen, and even use them as inline ref, whereas the 1' source is perfectly legitimate for this purpose because as a quasi-verbatim citation no OR is involved.
I should perhaps phrased it differently: "Introduce orig. descr. as the standard inline ref for taxobox author/date info".
3. Regarding use of primary sources. Some opt for a very restrictive policy here. However, this would mean speedy-deleting most of Category:Fossil taxa described in 2011 and uber pissing off WP:PAL and WP:DINO. Whether these articles, though by necessity relying on a single 1' source, violate WP:SCHOLARSHIP, WP:PSTS, WP:RS, WP:NOR etc in any way whatsoever is very much arguable, I'd argue. As a rule, these articles only state with certainty what is certainly known, and any statements beyond are habitually disclaimered ("may, include, at least, requires further research, still undetermined"...), and aim to provide secondary sources indirectly pertinent to the matter (e.g. info in the lagerstätte) and thus fulfil the #1 rule of WP:V: WP:CHALLENGE. (Theoretically, not using "is a proposed [species, genus...]..." violates NOR though. This is in fact significant when it comes to nomenclatorial disputes.)
4. Restated for clarity: my remarks above are concerned with the proliferation of internal nomenclatorial inconsistencies. For taxonomy, secondary sources are usually available (but see above).
Go back to the Theritas example above. Compare:
Theritas paupera (C. & R. Felder, 1865)
Theritas danaus (C. & R. Felder, [1865])
How to determine which is correct by any other way than consulting the 1' source and applying the ICxN rules? I bet you we will find at least 1, probably 2 more variants even in the usually-reliable databases, if any exist on this genus. For the Novara expedition, we might actually get a secondary source (scholarly, non-peer-reviewed, e.g. zoonomen). For less monumental 1' sources this is not the case. Dysmorodrepanis (talk) 20:07, 1 April 2012 (UTC)
I don't know what kind of discussion this is supposed to engender, but it reads like a setting-out of one person's opinion, rather than an attempt to develop a consensus. I don't wish to sound mean, but essays such as this probably belong in user-space, not here. --Stemonitis (talk) 20:19, 1 April 2012 (UTC)
Well I think that the fact that we have a large number of genus pages where species are listed with author/date info and no parentheses, yet were describes before the genus -- whereas we had few of these 2 years ago -or so -, is indicative of a general quality-control problem and declining editorial standards of WP:TOL, and would like to know if anyone else has noted. Dysmorodrepanis (talk) 20:31, 1 April 2012 (UTC)
I see. I wouldn't have guessed that from your previous post. Do you have anything beyond anecdotal evidence that things are getting worse? Can you establish who is doing it, and where the errors are coming from? It would be good to discuss the actual problem, rather than trying to deal with it through general assertions about primary and secondary references, and so on. --Stemonitis (talk) 21:47, 1 April 2012 (UTC)
I, too, have noticed no such increase in author citation errors. In fact, I think our collective efforts have only served to make it much more reliable, one edit at a time. Again, I don't think this a special case with respect to author citations. Existing guidelines and policies cover these subjects. If you notice errors, WP:SOFIXIT. If you think (and can present relevant examples) that a specific database is more error prone than others, we can discuss a suggestion to not use that database. Even then, though, we could just write a short guideline that says editors should try to verify author citations for that taxon in multiple reliable sources, but again I fail to see how this is different or more important than any other information in an article. I'd also like to note that the threshold for inclusion in Wikipedia is verifiability, not truth. The databases and other reliable sources used to obtain data on author citations for taxa reach that threshold. I see no problem. Rkitko (talk) 22:23, 1 April 2012 (UTC)
The article muntjac states that the subfamily of the genus Muntiacus is Cervinae, while Indian muntjac says that it is Muntiacinae. Apparently, both have been used; see [2]. Which one are we going to go with, though? It is now inconsistent across articles. InverseHypercube (talk) 07:03, 10 April 2012 (UTC)
Mammal Species of the World explains: "Muntiacini generally has been regarded as a subfamily (Haltenorth, 1963) and has usually been attributed to Pocock, 1923, Proc. Zool. Soc. Lond., 1923:207; treated as a full family by Groves and Grubb (1990); relegated to tribal status in Cervinae, by Groves and Grubb (1987) and Grubb (2000b) supported by evidence in Kraus and Miyamoto (1991)." --Stemonitis (talk) 07:12, 10 April 2012 (UTC)
Ah, okay. Thank you. I'll change all the muntjac articles to use this classification. InverseHypercube (talk) 15:37, 10 April 2012 (UTC)
Article titles for monotypic taxa
Hello everyone. I wanted to ask whether the Tree of Life project has an agreed-upon convention as to which name to use as the title of an article about a monotypic taxon? For example, do we use the binomial name of the species as the title of an article about a monotypic genus? Or do we use the name of the genus as the title? Thanks, Invertzoo (talk) 15:58, 10 April 2012 (UTC)
We use the genus, unless that title's ambiguous. If the genus name is ambiguous (e.g. "Gregoria"), we use the binomen (e.g.Gregoria fenestrata), unless that is also ambiguous (rare, but it could happen). --Stemonitis (talk) 16:04, 10 April 2012 (UTC)
It's sort of an article. Sort of a dab page. Could someone please look at it and push it one way or the other? Many thanks. Anna Frodesiak (talk) 23:47, 23 May 2012 (UTC)
List of animal cartoon characters sorted by biological taxonomy
I am interested in seeing an article "List of animal cartoon characters sorted by biological taxonomy".
—Wavelength (talk) 19:00, 29 June 2012 (UTC)
Two articles with the same taxobox
A taxobox was added to the article pet skunk. The use of this animal in this way is a subset of the taxon Mephitis mephitis, and as such shouldn't get the same taxon, don't you think? Chrisrus (talk) 05:16, 5 July 2012 (UTC)
I've removed it. From sources, it's not restricted to M. mephitis at all, although the latter is the most commonly domesticated.-- OBSIDIAN†SOUL 08:24, 5 July 2012 (UTC)
Discussion at Wikipedia:Categories_for_discussion/Log/2012_July_5
You are invited to join the discussion at Wikipedia:Categories_for_discussion/Log/2012_July_5. Several bird categories have been nominated for renaming because their latin names are ambiguous. Inputs welcome. KarlB (talk) 18:59, 9 July 2012 (UTC)
Heads up......
Any big/broad biology articles that are in poor shape are fair game for Wikipedia:Core_contest over the month of August? WWII might be a tad tricky but surely there are some other big'uns.....Casliber (talk·contribs) 00:20, 18 July 2012 (UTC)
Darwin was Nearly Right
Off-topic discussion of work by Eugene McCarthy
We all know that taxonomy is a 'bag of worms' and that a species may be (and often is) classified with virtually equal accuracy from a dozen different perspectives. Ever stopped to wonder why?
Darwin was spot on with natural selection, and so very close on many occasions to the real source of genetic variation. This approach by McCarthy unlocks the last riddle - Saltation - which plagued Darwin's theory.
Unfortunately, it demonstrates that the other half of the 'Tree of Life' is missing - there is now a huge amount of work to be done to identify 'the other half' and start to put in place all the crossing links.
There's a "brief" synopsis here for those of us who are TLDR-challenged. AFAICT, he is basically restating the well-known genetics behind punctuated equilibrium, although I'm not sure he realizes that. He's not the first to challenge the gradualism of Darwin's views, and if investigators continue to ignore the older literature, he won't be the last.--Curtis Clark (talk) 16:16, 15 June 2012 (UTC)
It seems to start off OK (e.g. what on a cursory skim looks like a decent review of hybridisation), but gets worse (e.g. armadillos are descended from ankylosaurs, and whales from mososaurs). Lavateraguy (talk) 16:20, 15 June 2012 (UTC)
Yes, I'm afraid it gets solidly into very shaky ground there. When you put their plates flat on their backs, stegosaurs look kind of like pangolins, so pangolins must be surviving stegosaurs—never mind the fact that stegosaur plates are bony, while pangolin ones are epidermal (quite apart from the numerous other features that identify one group as mammalian, the other as archosaurian). And I wonder what made him think that multituberculate and rodent dentitions are similar. Ucucha (talk) 17:51, 15 June 2012 (UTC)
@Ucucha - you are thinking in classical Neo-Darwinian linear evolutionary terms with this response - with hybridisation characteristics of each parent are shared / mixed up making it entirely possible for bone plates to become epidermal in the hybrid. We don't understand the rules of hybridisation yet - when do hybrids come out half and half, when do they come out blends, but the old way of thinking stands in the way of 'seeing' what seems to be in front of our noses. DerekSmith (talk) 20:16, 15 June 2012 (UTC)
If we don't understand the rules of hybridization after all this time, we must not understand much of anything. Don't get me wrong; I even go looking for hybrids. But there's no revolutionary new science here.--Curtis Clark (talk) 01:53, 16 June 2012 (UTC)
Well, Multituberculata does say "multituberculates had a cranial and dental anatomy similar to rodents" ...
I'd guess that he doesn't know anatomy and palaeontology, so there's nothing to stop him running with impressionistic similarities. The lack of DNA from fossil organisms shields his conjectures from DNA testing to a degree, but I expect that sequence data doesn't match the expectations of his conjectures. Lavateraguy (talk) 18:09, 15 June 2012 (UTC)
Snakes are descended from annelids! —innotata 19:59, 15 June 2012 (UTC)
Some of the suggestions may well to turn out to be wrong, but what of the principle that every 'species' did not slowly evolved through gradual mutation, but sprang into (geologically) instant existance through hybridisation of two compatable parents. It happens frequently in ducks and fish, sometimes the parents are driven out because of the hybrid vigor and environmental suitability of the hybrids and a new species is born. Does this not explain the extreme problems often encountered in taxonomy - each 'species' does not have one parent, but two, and there are no 'missing intermediary fossils' because there were none. DerekSmith (talk) 18:52, 15 June 2012 (UTC)
The simple answer is "no". Some species arose by hybridisation (it's quite common in plants), but DNA sequence data generally supports a dichotomising tree of descent. If wide hybridisation was a common occurrence it would show up as more horizontal tranfer in genomes than is seen. (BTW, can you give even one example in ducks - ducks, geese and swans hybridise readily, but I'm not aware that this has resulted in a new hybridogenetic species. Outside ducks, there is an example in Passer.) Lavateraguy (talk) 19:12, 15 June 2012 (UTC)
Interspecific hybrids are fairly common, intergeneric hybrids are generally rare, but common in a few groups (e.g. orchids, cacti, pome fruits, water fowl, wild fowl) and not all that rare in general in birds and plants, and interfamilial hybrids very scarce (all that I recall are among wild fowl); I get the impression that he is talking about rather wider hybridisation. Lavateraguy (talk) 19:12, 15 June 2012 (UTC)
There's rather shaky grounds to say even the Italian Sparrow or Mariana Mallard are hybridogenetic. Genera and families are of course arbitrary, but I doubt there's any interfamilial hybridisation under modern taxonomies (there are only two families of wildfowl, Magpie Geese and everything else). —innotata 19:59, 15 June 2012 (UTC)
Yes, I'm remembering the galliforms now, though don't know how they fall in newer classifications. (Wildfowl is the British term for waterfowl, galliforms are called landfowl or gamefowl.) —innotata 20:18, 15 June 2012 (UTC)
[ec] When I still did research, I studied hybrid speciation in plants. One of the consequences of genetics is that hybrid species ordinarily share apomorphies of both parents. In plant groups with a fair amount of hybrid speciation, divergent speciation (as measured by patterns of apomorphies) greatly outweighs hybrid speciation (in the genus I studied, Encelia, only two of over a dozen species had well-substantiated hybrid origins. (These were diploid species, not alloploids.)
I'm responding quickly here, but I can look up studies that state that the vast majority of angiosperms (and therefore the vast majority of plants) are of hybrid origin. So a genus having just two members of hybrid origin does not strike me as a plant group "with a fair amount of hybrid speciation." But, of course, the mere fact that the other members of Encelia were not shown to be of hybrid origin does not mean they aren't. Nor does it mean that they are of gradual, nonhybrid origin. The tendency, all too often, I find, is to ASSUME gradual origins until hybrid origins are firmly demonstrated. In the present context, that's a real bias. Koolokamba (talk) 17:25, 16 June 2012 (UTC)
On the other hand, any character state distribution can be explained by hybridization events; i.e., hybridization cannot be falsified through phylogenetic analysis. In the final edition of Plant Speciation, Verne Grant made the statement that hybridization in flowering plants was so pervasive that we would never figure out their phylogeny. It didn't turn out that way.--Curtis Clark (talk) 19:30, 15 June 2012 (UTC)
Was that before DNA sequencing became technically and economically feasible? Lavateraguy (talk) 20:10, 15 June 2012 (UTC)
In my specific case, we found chimeric ITS sequences and shared RAPD markers. But the Angiosperm Phylogeny Group demonstrates that congruent molecular and morphological trees are reasonably common.--Curtis Clark (talk) 01:50, 16 June 2012 (UTC)
The intended target of my question was the statement made by Verne Grant. Lavateraguy (talk) 13:49, 16 June 2012 (UTC)
Any character state can be explained, that is, a phylogeny can be constructed, also under the assumption that no hybridization has occurred. But that doesn't make the explanation true. So, yes, "hybridization cannot be falsified through phylogenetic analysis," if by phylogenetic analysis you mean "construction of a phylogeny." But if you look at the situation historically, you do see a pervasive phenomenon across a broad range of taxa: inconsistent phylogenies have been produced by different research groups when different sets of traits/genes were used as a basis of comparison. In this respect, phylogenetic analysis speaks clearly for a pervasive role of hybridization in evolution. And, I think it's fair to say that it also suggests that the current, most popular angiosperm phylogeny is but the latest most popular angiosperm phylogeny. There have been many others, in many ways inconsistent with the current one, and if we keep thinking the same way, imagining a true tree must exists when everything is actually reticulate, this historical trend will, no doubt, continue. Before long we'll have another most popular angiosperm phylogeny. But isn't that just a bit sisyphean? Looking at things historically, people engaged in the production of phylogenies have always claimed that their own phylogenies are valid and satisfactory (how could they not?), and yet in each case subsequent workers using new data sets come up with new, supposedly more accurate phylogenies. But the mere continuation of debate here suggests that different data sets are generating different phylogenies (the expectation under the hypothesis that hybridization is rampant). I think this is what McCarthy is arguing. — Preceding unsigned comment added by Koolokamba (talk • contribs) 17:39, 16 June 2012 (UTC)
Didn't know he was the same person. I've read it (and cited it on Wikipedia), and don't remember any suggestion of large-scale hybrid descent in its discussion or from its contents. —innotata 20:18, 15 June 2012 (UTC)
You've misinterpreted it. It claims (and I believe it) hybrids with 4,000 different species parentages. A hybrid (nothospecies) is not the same as the species. For example ligers and tigons, reciprocal crosses between Panthera leo and Panthera tigris are not (yet, and probably will never be) a hybridogenetic species.
If you were to read the book you'd find that something like 90% of those pairings represent intrageneric hybrids (between a natural tendency to broader hybridisation, taxonomic biases, and human intervention birds have a relatively high rate of intergeneric hybridisation).
For an accessible database of avian hybrids see hereLavateraguy (talk) 20:20, 15 June 2012 (UTC)
Thanks, my mistake, as I said, I had not read the book and misread the summary, indeed, it says thousands of hybrids, not species. Would it be reasonable to think though, that, in the fullness of time, some of those hybrids might establish themselves as species? Wouldn't this be a demonstration of the McCarthy proposal? DerekSmith (talk) 21:02, 15 June 2012 (UTC)
Thanks for posting it, hadn't seen it before. Before anybody says it supports McCarthy's claims, note that the site doesn't distinguish between copulation and actual (much less fertile) hybrids—it's a database of sources to look at. —innotata 20:28, 15 June 2012 (UTC)
There are two parts to the proposal, first that hybridisation creates the instant genetic difversity, then, second, Stabilisation Theory under natural selection fixes the genetic makeup and increases fertility. Massively reduced fertility is initially compensated for by improved fitness. DerekSmith (talk) 21:07, 15 June 2012 (UTC)
Just to be fair to McCarthy, guys, can any one cite even one instance where a species is KNOWN to be of gradual, not hybrid origin? Granted that a very small percentage of species are KNOWN to be of hybrid origin. But these are documented. Koolokamba (talk) 21:25, 15 June 2012 (UTC)
Any autopolyploid species. But, if you want to bend over backwards, novel autopolyploid populations may also display genomic shock. Lavateraguy (talk) 22:19, 15 June 2012 (UTC)
As a point of order, McCarthy self-published this "book" on his website, so it's a WP:SPS. I haven't evaluated his resume to see if he's an expert in his field, but one does wonder why he didn't get it published by one of the many academic press operations out there. Rkitko (talk) 21:31, 15 June 2012 (UTC)
"Eugene M. McCarthy earned his Ph.D. in genetics from the University of Georgia, where he has conducted research into genomics and hybridization for more than a decade." He has a few papers on LTRs, and on old paper on recombinational speciation. Lavateraguy (talk) 22:17, 15 June 2012 (UTC)
Thanks. Of his own admission, the book was self-published because he couldn't get it published elsewhere. He claims reviewers were just cruel and clearly blind, which lead to constant rejection. I can sympathize with repeated rejection, but there is almost always a good reason. I'm leaning toward an assessment of the work as fringe, at the moment. Rkitko (talk) 23:05, 16 June 2012 (UTC)
I'm at a library, guys, and I see that in the introduction to his Avian Hybrids book McCarthy lists (pp. 29-30) a lot of birds of hybrid origin that have been treated as species. I also see, in flipping through the book, that most of the crosses seem to be interspecific, not intergeneric as somebody claimed above. 72.152.228.149 (talk) 21:44, 15 June 2012 (UTC)
"intrageneric" rather than "intergeneric" - interspecific includes intergeneric and interfamilial, intrageneric excludes them. Lavateraguy (talk) 21:51, 15 June 2012 (UTC)
Seems to me that a hypothesis of pervasive hybridization COULD be falsified: If it were true for the typical set of related organisms, then if distinct gene trees were concordant, it would be consistent with the idea that hybridization was rare. If, on the other hand, different genes regularly generated distinct species phylogenies, then the pervasive hybridization would be supported. I won't name any names, but a GREAT figure in population genetics once told me that the only way to get clean results with phylogenies is to limit the number of markers. In simple terms, his advice was don't look at too many genes or you'll get conflicting results and you won't be able to publish. 72.152.228.149 (talk) 22:01, 15 June 2012 (UTC)
In repsonse to Genomic Shock's comment above: An autopolyploid is the product of a spontaneous chromosome doubling that generally occurs in ONE generation--not a very good example of gradual origin if you ask me. Actually, I see autopolyploidy is one of the stabilization processes that McCarthy mentions on his site. Koolokamba (talk) 22:13, 15 June 2012 (UTC)
I did a site search of McCarthy's site for the term "autopolyploid" and on one page he writes this: "Many supposed autopolyploid [sic] are actually derived from hybridization, but are not recognized as such due to the official taxonomic treatment of the hybridizing forms. That is, they are products of hybridization between forms that are distinct, but treated as conspecific. Suomalainen et al. (1987: 100) state that polyploid vertebrates of known origin have generally 'proven either to be species hybrids or hybrids between different cytological races [i.e., different chromosets] of a single species.' " Koolokamba (talk) 22:21, 15 June 2012 (UTC)
What you meant by gradual was unclear, especially as you opposed it to hybrid. Lavateraguy (talk) 13:45, 16 June 2012 (UTC)
Stebbins called these "segmental allopolyploids" in plants. Many botanists dont' regard strict autopolyploidy as a means of speciation, since there is continued gene flow into the polyploids through non-reduced gametes.--Curtis Clark (talk) 01:45, 16 June 2012 (UTC)
Taking the comment above "GREAT figure in population genetics once told me that the only way to get clean results with phylogenies is to limit the number of markers. In simple terms, his advice was don't look at too many genes or you'll get conflicting results and you won't be able to publish. 72.152.228.149 (talk)" -- isn't this true across the board? and if so, does this not directly imply that hybridisation is behind the vast majority of species origins? And if this is true, then is there not a huge amount of taxanomic work to be done to identify the other hybridisation parent for each of these hybridisation generated species? The 'simple' linear 'Tree of Life' then takes on the form of a genealogical structure with most of the crosses being quite close, but some conceivably quite distant if the Hybridisation Zone was stable and the hybridisation compatability high enough.
DerekSmith (talk) 12:25, 16 June 2012 (UTC)
Common descent with dichotomising speciation results in nested hierarchies inferred from different data sets being correlated - not identical. It's generally better to improve taxon sampling that to throw base pairs at a taxonomy - more loci means more opportunities to find a long branch artefact. On a smaller scale, differential fixation of ancestral polymorphisms results in differences between the trees inferred from different loci.
Even if you got more greatly divergent trees this wouldn't imply that hybridisation was behind the vast majority of species origins - if one species in 10 resulted from hybridisation between divergent parents that would be enough to mess up the trees. Lavateraguy (talk) 13:42, 16 June 2012 (UTC)
This reasoning begs the question. If descent truly were strictly dichotomous, then it would be fine to talk about correlated data sets. However, given the two hypotheses we are considering (A: descent is dichotomous and hybridization so rare it can be ignored; and B: descent is reticulate and hybridization is pervasive) any contradictory data that reduces the correlation is actually evidence for hybridization and cannot be merely discounted (as it usually, and reasonably, is under the assumption that hybridization is rare). The comment about differential fixation reflects the same bias: it assumes that the differences are due to differential fixation when they might just as well be evidence of hybridization.DerekSmith (talk) 16:33, 16 June 2012 (UTC)
So, this is a nice discussion, but is there any point to it? Are you suggesting improvements to an existing article or trying to gain support for a policy or guideline, or alert other editors to an existing issue in an article? Those are the typical functions of a Wikipedia talk page. All I see here is so far is a decent discussion of the article you posted, its merits, and the ideas within that article. I see no connection to how it is to be used on Wikipedia. There are better venues to continue that discussion as Wikipedia is not a forum. Have any specific proposals on what you want to see done? Or was this just a general "for your interest" post? If the latter, I suggest wrapping up the conversation. What is clear from this discussion is that many of our articles on phylogenetic topics could use improvement. For example, at present incomplete lineage sorting currently redirects to coalescent theory. Rkitko (talk) 17:26, 16 June 2012 (UTC)
Seems to me there's a VERY BIG POINT. This discussion has a direct bearing on how the "tree of life" should be treated in this article: Is it (i.e., the tree of life) a valid notion or not? If it probably isn't, SURELY that should be at least mentioned in the article. Likewise, if there is disagreement concerning its validity, that should be mentioned, too. — Preceding unsigned comment added by Koolokamba (talk • contribs) 18:47, 16 June 2012 (UTC)
Good point Rkitko, I was starting to worry that this discussion was getting quite large. Is there any point to it? Again good question. From the discussions so far posted, it seems that there is a significant probability that the two processes proposed by McCarthy are substantiated, and that they answer the Saltation problem as yet unanswered by Neo-Darwinism. Consequently the subject of this page - WikiProject Tree of Life will be found to be sadly lacking in describing the lineage of species. Had McCarthy's book been trashed by the expert editors here present, that would have been the end of it, but it wasn't, in fact some posters brought forward supportive examples. So rather than asking is there a point to the discussion, it would perhaps be more pertinent to ask - how are the editors going to correct the present rendition of this page to adapt to new science? DerekSmith (talk) 18:57, 16 June 2012 (UTC)
Derek: Beyond the scope of whether we think there's that much new science presented, its merits, and whether we think it's revolutionary (I don't, and frankly I'm beginning to think much the same that Peter coxhead said below), Wikipedia does not respond to immediate changes like this. McCarthy has proposed ideas. Have others opined on the weight of his evidence? Have others supported it? Has it been incorporated into phylogenetics textbooks? Wikipedia is descriptive, not prescriptive. We will incorporate McCarthy's ideas once the relevant fields have done so. That is the nature of an encyclopedia. Rkitko (talk) 23:05, 16 June 2012 (UTC)
The claim that in a high proportion of cases speciation in eukaryotes proceeds via hybridization is an extraordinary one and would require extraordinarily good evidence. It's at odds with the vast body of molecular phylogenetic data accumulated since the mid-1990s and seems to be sourceable to a single self-published book. Classic fringe science as far as I can see. So there is at present no need to make any changes. Peter coxhead (talk) 19:19, 16 June 2012 (UTC)
Cite your sources, please, not your opinions. "Vast body" says nothing. And please keep in mind it's not a "claim." It's a hypothesis with a great deal of empirical support. Angiosperms, for example, are eukaryotes, and, as almost any biologist must have heard by now, most angiosperms are of hybrid origin. Koolokamba (talk) 19:40, 16 June 2012 (UTC)
Cite your sources that "most angiosperms are of hybrid origin".--Curtis Clark (talk) 23:12, 16 June 2012 (UTC)
Yes, please. Let's avoid ad hominem arguments! Sylvia 1024 (talk) 21:03, 16 June 2012 (UTC)
Peter, your point about there being a 'vast body of molecular phylogenetic data' is aptly at the very heart of this discussion, because as pointed out above by 72.152.228.149, (s)he was advised by a 'GREAT' name that this means of grouping is so confused that the researcher has to be utterly blinkered in order to make any connection at all - vis - "the only way to get a clean result with phylogenies is to limit the number of markers". Indeed, McCarthy makes exactly the same comment himself, and goes on to explain the reason for this being due to the fact that the hybrids contain mixed markers from both (all) parents of the cross. A vast quantity of 'manipulated' data is still manipulated data. Perhaps it is time that the editors of this page should look to amending the phylogenies of at least the 'species' now known to be hybrids, and then develop the page from there. Could we perhaps start by developing a consensus of known hybrids? DerekSmith (talk) 23:15, 16 June 2012 (UTC)
The book in question is still self-published, and the anatomy and palaeontology is still shaky, to be generous. —innotata 23:11, 16 June 2012 (UTC)
A comment by an unnamed person reported by an anonymous person isn't evidence for anything. Molecular phylogenetic work published in the late 1990s and early 2000s did use a relatively small portion of the genome. More recent work uses "multigene supermatrices" and/or compares trees produced from different subsets of sites within genomes and the criticism simply doesn't apply. (A couple of examples I've read recently are: Qiu, Yin-Long (2006), "The deepest divergences in land plants inferred from phylogenomic evidence", Proceedings of the National Academy of Sciences, 103 (42): 15511–15516, doi:10.1073/pnas.0603335103((citation)): Unknown parameter |coauthors= ignored (|author= suggested) (help) and Zhong, Bojian (2011), "Systematic error in seed plant phylogenomics", Genome Biology and Evolution, doi:10.1093/gbe/evr105((citation)): Unknown parameter |coauthors= ignored (|author= suggested) (help).) Similarly, early work used rather simplistic and atheoretical methods of analysis, such as parsimony. More recent work uses models for genomic changes which can be tested statistically. I'm not saying that there aren't still some papers published based on carefully selected sites, but there are now many that are scientifically much stronger.
These molecular phylogenetic approaches have clearly shown when the splitting/branching model fails, e.g. because lateral gene transfer has taken place (e.g. between some angiosperm parasite species and their hosts). Equally they have clearly shown that the branching model is correct in most cases. (At least as regards the rarity of hybridization across widely separated branches. In groups where significant gene flow is still occurring between populations morphologically recognized as "species" (e.g. Ophrys, Dactylorhiza), it doesn't make a lot of sense genetically to talk about "hybrids between species".) Peter coxhead (talk) 07:26, 18 June 2012 (UTC)
One thing to note. If species A splits into species B and C, and species B and C hybridise to produce species D, then tens of million years later, if the only descendants of A are also descendants of D, then the fact of species hybridogensis in the lineage doesn't change the tree of living species.
Reticulation in various forms is widespread, and is one of many processes that can interfere with producing a clean tree from DNA sequence data (see also parology, base substitution biases, variations in base substitution rate, homoplasy, ...). Much of what McCarthy writes is not novel, and is not ignored by phylogeneticists (and doesn't require modification to the tree of life in Wikipedia). Lavateraguy (talk) 08:28, 18 June 2012 (UTC)
Arbitrary break to make editing easier
Derek, I assume you mean a "consensus of species known to be of hybrid origin". A list of known hybrids would be more noise than signal here, except to someone who already had their mind made up about mechanism.
Incongruent phylogenetic data can be explained as homoplasy or as lateral transfer (through hybridization or other means). Any homoplasy can be explained as lateral transfer, and any lateral transfer can be explained as homoplasy. So appeals to incongruence are not useful in convincing people, since it can mean anything.
If I'm understanding the mechanisms he postulates, at least some of them are well-known and documented (recombinational speciation, for example). He's saying nothing new, at least there, about mechanism; it seems to me that his new argument is about scope, and the burden of proof is on him. There are ways to positively identify species of hybrid origin, but afaik they still require examining each candidate species individually.--Curtis Clark (talk) 00:40, 17 June 2012 (UTC)
On his website, McCarthy quotes John Avise (someone, I trust, all the phylogeneticists among you will have heard of) as follows about his, McCarthy's, theory:
"In his recent book, In the Light of Evolution (2008), renowned evolutionary biologist John Avise says that if the views presented on this website are correct, our ideas of biology and the evolutionary process will have to be revised at a fundamental level. There, he [i.e., Avise] states that: 'First, phylogeneticists would have to admit that their dream of reconstructing a branched tree of life had been merely a pipedream, and they would have to accept the new and probably far more difficult challenge of working out the precise history of reticulation events for each organismal group and how such reticulate genealogical histories have idiosyncratically distributed particular bits and pieces of DNA from disparate sources to extant taxa. Traditional concepts of species, phylogeny, ancestry, and classification, as well as the significance of reproductive isolation, would all have to be reevaluated. Biologists would have to embrace the notion that biological processes falling somewhat outside the standard neo-Darwinian paradigm for speciation (such as interspecific hybridization and the reproductive stabilization of genetic-recombinant derivatives) could play major and previously underappreciated roles in evolution. They would have to reevaluate the origins of genetic variation on which natural selection acts and how novel phenotypic adaptations and different forms of life mechanistically come into being. In short, major shifts in evolutionary thought would be required, and this would open wonderful opportunities for the eventual emergence of a grandly updated evolutionary synthesis, 21st-century style.' — From In the Light of Evolution, Volume II: Biodiversity and Extinction, 2008, pp. 288–289." McCarthy took the quotation from Avise's online discussion of the theory (Avise calls it the "Network Theory"), which is available on line . Here's the link: http://books.nap.edu/openbook.php?record_id=12501&page=288 Koolokamba (talk) 22:21, 17 June 2012 (UTC)
So... one favorable mention in 2008 from someone McCarthy once worked with at the University of Georgia (probably the only reason he was aware of the work) and then crickets since then? I'm sorry if this sounds a bit harsh, but I don't think this discussion is productive any longer. McCarthy didn't even address one of the sources of homoplasy, incomplete lineage sorting, in his work. One would think that such important ideas would at least warrant a mention, especially if he's suggesting it is not as important as hybridization. The point? It does not seem like the important paradigm-shifting work that McCarthy, as a self-promoter, and his supporters here want it to be. There is nothing of relevance to Wikipedia that can come from continuing, unless we collaborate to work on some of the articles that are in desperate need of attention that I pointed out above. And continued promotion of McCarthy's ideas should not be tolerated until they gain wider acceptance - see WP:NOTSOAPBOX and again WP:NOTFORUM. Rkitko (talk) 23:00, 17 June 2012 (UTC)
Collaboration entails entertaining alternative viewpoints. I don't know about anyone else who's looking at this conversation, but I can't see much of that here. So I, for one, will leave you to your phony, pointless tree of life and move on to other articles, NOT suggested by you, narrow-minded Rkitco, Mr. Keeper of the Truth. And I'm really NOT sorry if that sounds a bit harsh. Koolokamba (talk) 23:23, 17 June 2012 (UTC)
Actually, when I went away, I was copyediting the article on the Moghul Empire, and there I noticed alternative viewpoints are in fact allowed, for example, alternative views on the reasons for the empire's decline during the 18th century. So it really isn't true that unestablished alternative views aren't allowed on Wikipedia. It's that you, Mr. Rkitco, are setting yourself up a little fiefdom here where you can suppress opinions that differ from your own. You don't even have your Ph.D. yet. So how can you possibly in good conscience set yourself up as such an expert? Even someone with a degree shouldn't do that. Anyway, bye. This time for good. Koolokamba (talk) 23:53, 17 June 2012 (UTC)
(edit conflict)Collaboration in building the encyclopedia certainly does include entertaining alternative views, but we as editors do not decide which views have merit. We simply describe the current consensus of whatever field or topic we're writing about and do so with a neutral point of view. Yes, this also includes providing information on major competing hypotheses, but not WP:FRINGE ones and certainly not ones that have not been fully evaluated by other scientists and authors. What I see here is anything but and is extremely thin on details on what the proponents want us to do with McCarthy's self-published book. I don't see, given how little attention has been paid to his ideas, how we could incorporate anything from that manuscript into Wikipedia. Also, what you observe as narrow-mindedness is not; I am simply reminding you (or introducing you) to what Wikipedia is not. Please avoid personal attacks of that nature. My qualifications are not of concern, here, and I provide them for those that are curious. I did not claim to be an expert on the subject, but I did evaluate the manuscript in terms of its merits as a reliable source for use on Wikipedia. That's what matters here. Please stick to that discussion and do not get distracted by my personal details. Cheers, Rkitko (talk) 00:05, 18 June 2012 (UTC)
LOL. GOODBYE, MR. PRONOUNCER OF THE CONSENSUS. Koolokamba (talk) 00:16, 18 June 2012 (UTC)
Oh, and folks, unless there was any mistake, it is possible that Koolokamba (talk·contribs) is indeed Eugene M. McCarthy himself, here to promote his own work. On a website that linked to his own, someone with the same username, koolokamba, commented and signed it "Gene McCarthy Ph.D. Genetics" after thanking the original poster for linking to his website. So there is an added conflict of interest and I wouldn't be surprised if the WP:SPA Sylvia 1024 (talk ·contribs) who jumped into this discussion and edited twice turned out to be a sockpuppet or meatpuppet. Rkitko (talk) 00:20, 18 June 2012 (UTC)
That's just sad, that someone with actual academic credentials should fall so low that they would feel compelled to defend their ideas on Wikipedia. --Curtis Clark (talk) 03:20, 18 June 2012 (UTC)
Yes, I freely admit that Koolokamba is Gene McCarthy. And if I was deceptive in referring to myself in posts in the third person, then I apologize for that. But it wasn't my idea to post anything on this page in the first place. That was Derek's, and he started posting without telling me he was going to do so. I learned of what he was doing only after he'd been doing it for a while and sent me an email. I had even advised him specifically against approaching phylogeneticists with my ideas. I'm sorry, but what I told him was that phylogeneticists, from my own experience, are the least likely of any class of biologists to embrace my account of the evolutionary process. After all, if descent is not at least generally dichotomous, then they are out of business. But once I went to this page and saw him fighting alone against you, without the twenty-five year experience that I've had in your arguments, frankly, I felt sorry seeing you gang up on him. So I started to comment. You repudiate me because I'm "self-promoting." But what is any scientist today? Isn't every seminar, every talk at every retreat and conference, an exercise in self promotion? Many of the most respectable scientists I know, seem never to let up with promoting themselves, they extend it into their lab meetings, the classes they teach, and even go on and on when you go out to drink with them. So I really think it's really kind of a joke to me that you call ME self-promoting. Until Derek, who's a retired microbiologist, by the way, contacted me a few months ago I had been working strictly on a novel for more than a year. So not much self-promotion there. Derek though, is so enthusiastic about the overall theory that I present in my book, that we've exchanged a lot of emails lately with him questioning me about my theory. That's the only thing that's gotten me off my intellectual duff. So I believe my posts to this page are my first efforts to promote this book in more than two years. Really, other than making it available on my website, I've done very little to promote it for even longer than that. My last real effort on that front was in responding to reviewers when it was under contract at Oxford University Press. Those reviews, of which as I recall there were five or six, ran the full spectrum from glowing to damning. The most unfavorable comments in the reviews, however, were always to the effect that "this contradicts accepted thought." Well, of course, any new theory contradicts accepted thought. So I refused to make changes on the basis of such criticism, and after a year of arguing with me, Oxford terminated the contract. Not wanting to spend another fruitless year with another publisher, and also thinking the prospects for an open-minded reception elsewhere were poor when my submission had raised such a storm at Oxford (where I had successfully published before), I simply made the book available on line and moved on to other things. That was how John Avise saw it, by the way. Although he was on my Ph.D. committee, and we're on a first name basis, I haven't seen him for eight or nine years, and I've only spoken to him once when I called him up to thank him for his comments on my book. So you wouldn't exactly call him my friend. But I'm going on far too long. Before I close, though, I should say I'm sorry for my bitter words in some of my last posts, I was tired at the time and I didn't like being called "fringe" or being talked down to since I hold a B.S. in mathematics, a Masters in genetics, and a Ph.D. in genetics at the University of Georgia (which is consistently rated in the top ten genetics departments in the country). My career in genetics began almost twenty-five years ago, and I wouldn't be surprised if I have more peer-reviewed publications in genetics and biology than anyone else posting to this page. So that's not fringe. And if it's become fringe to propose a new theory in biology today, then we've got a real problem. And is it really new? The only way you can make an honest decision about that is to read what I have to say--from beginning to end. Skipping around and picking out ideas and taking potshots at them out of context, and gossiping in a forum will never get you there because, as I say, a new theory is always going to contradict things you think are right if your thought is laden with another theory. If you give me a fair hearing, I think you'll see that the system of thought that I present will allow you to understand many biological phenomena that have puzzled you under conventional theory. I do understand, though, that an encyclopedia is not the best place to introduce new ideas. (I wouldn't be here if it weren't for Derek.) But I do think, that if you will take a few hours to read what I say in my self-published book, that at least some of you will reconsider the idea of spending much of your time working on the "tree of life." Koolokamba (talk) 14:11, 18 June 2012 (UTC)
I just have to respond to the allegation that all scientists self-promote: real scientists concentrate on research; don't justify your self-aggrandizement by tarring others. Promoting our own publications in wikipedia is infra dig. Promoting a theory as "new" that is (1) a straw man fallacy and (2) a collection of other people's work, is similarly unworthy of a scientist. DS please also desist from spamming other editors' talk pages. Sminthopsis84 (talk) 20:57, 18 July 2012 (UTC)
Categories (section)
Added reference to Biota (ecology) in addition to the internal link Biota (taxonomy). The closed reference was placed at the end of the section as it was the first reference used in this article; being a lengthy article, uncertainty of MOS in this case the reference was placed in proximity to the reader's concern for ambiguity. O = MC4 22:45, 19 July 2012 (UTC)
How to make a cladogram?
Are there any guides how to write a cladogram for a Wikipedia article? FunkMonk (talk) 16:29, 20 July 2012 (UTC)
Thanks, I'll take a look. FunkMonk (talk) 17:32, 20 July 2012 (UTC)
I made an attempt at one here[3], but is there a way I can get the branch with Didunculus moved below the rest, instead of above? That's how it is in the paper. FunkMonk (talk) 20:42, 20 July 2012 (UTC)
Thanks, that's what I was thinking of, I tried something similar, but seems the placement of Didunculus should correspond exactly with the placement of its sister branch down in those. Thanks! FunkMonk (talk) 09:09, 21 July 2012 (UTC)
I've attempted to make a cladogram for moa, but the result is rather mangled: [4] Would anyone be able to fix it so it looks more like this?[5] The original figure is here[6], I have removed lineages within species from the other version. Thanks. FunkMonk (talk) 17:17, 14 August 2012 (UTC)
Have attempted a cladogram. Please check and modify as needed. The key is to add some dummy nodes. Shyamal (talk) 15:04, 15 August 2012 (UTC)
Thanks, looks correct. Is there a way to left align it? FunkMonk (talk) 15:12, 15 August 2012 (UTC)
Box moved to left. Hope that was what you meant. Shyamal (talk) 15:52, 15 August 2012 (UTC)
Exactly, and thanks again, I'll add "clear" so it won't interfere with the following sections. FunkMonk (talk) 16:00, 15 August 2012 (UTC)
Just a pointer to a discussion about creating an A-class review process for Tree of Life and related projects. - Dank (push to talk) 19:14, 23 July 2012 (UTC)
We're trying to give this article, NGSD, the right kind of infobox and otherwise find the proper place for it in our system.
Thanks for the link to the full pdf. I find that paper very unsettling. First, there's not a single taxonomist or systematist author. Second, all of the evidence they cite and discuss (more like a review than primary data) is largely anecdotal; there is very little data to go on. There are no statistical tests of the significance of the data. And clearly the authors have a POV to push since the lead author is associated with the NGSD conservation group (based in Oregon?). I tend to err on the side of caution as well and I would advocate, like they do, that this is an evolutionary significant unit for conservation purposes. But what that unit is called - a variety, subspecies, or species - is extremely unclear. Their argument for species status is not compelling. I may just be in my first year of a PhD program in taxonomy and systematics, but it's already clear to me that this paper would have required more data (molecular systematics, morphometrics) to be convincing. I'm surprised it was published. Regardless, I would be very careful when discussing it in the text or using it for citations. Avoid using it as the sole citation for an assertion, while still avoiding weasel words. Discuss it in the context of the uncertainty of the taxon.
As for the kind of infobox, at this point I see no reason to keep the taxobox on the article. The dogbox would be fine. Rkitko (talk) 14:24, 12 November 2011 (UTC)
Is that your opinion of the paper itself, or the references upon which the paper is based — or perhaps both? Specifically, the discussion at the NGSD article currently seems to revolve around the statement in THE NEW GUINEA SINGING DOG: ITS STATUS AND SCIENTIFIC IMPORTANCE at the end of page 47 and continuing on page 48: "It is, at minimum, an evolutionarily significant unit, and, at maximum, could be its own distinct species (Simonsen 1976; Gollan 1982; S Bulmer 2001; Bininda-Emonds 2002; Koler-Matznick et al. 2002; Clark et al. 2004; Savolainen et al. 2004). Therefore, for conservation purposes, the wild singing dog should be considered a unique taxon until the contrary is clearly demonstrated", which I used as justification for the "Unresolved" paramater in the taxobox. But it appears the issue is more complex than just relating to that parameter in the taxobox... Did you (or anyone else who understands this stuff) examine SIMONSEN V, 1976. Electrophoretic studies on the blood proteins of domestic dogs and other Canidae. Hereditas 82: 7–28[7] or SAVOLAINEN P, LEITNER T, WILTON A, MATISOO-SMITH E, LUNDEBERG J, 2004. A detailed picture of the origin of the Australian dingo, obtained from the study of mitochondrial DNA. Proceedings of the National Academy of Science USA 101: 12387-12390.[8] or CLARK LA, FAMULA TR and MURPHY KE, 2004. Evaluation of a rapid single multiplex microsatellite-based assay for use in forensic genetic investigations in dogs. American Journal of Veterinary Research 65:1446 – 1450.[9] (which seems to cost money to view) or any other of those eight references for that statement in the paper? If not, could you possibly take a look and give some feedback? I don't know if that information changes anything, but this stuff is beyond my layman's understanding of genetics. Mojoworker (talk) 22:15, 23 November 2011 (UTC)
Ok, I think that I can contribute to this. I read the first paper you referenced and it already listed the NGSD (Hallstromdog) as a seperate species and when you go by the differences in blood enzymes the jackal would be closer to the domestic dog. However this is at odds with the genetic data published in the second referenced article which clearly shows a connection with other dogs as well as the dingo. Another published article (http://www.nature.com/nature/journal/v464/n7290/full/nature08837.html) clearly showed that they are descended from wolves and among domestic dogs belong to the "Asian Group" together with dingoes, Chows, Akuitas and Shar-Peis. I also read a newly published scientifc article on the origin of the dingo, NGSD and Polynesian dogs (which can be found here: http://rspb.royalsocietypublishing.org/content/early/2011/09/06/rspb.2011.1395.full.pdf) and it states:
The study of mtDNA among Australian dingoes and Polynesian dogs showed that archaeological samples of pre-European dog from across Polynesia (the Cook Islands, Hawaii and New Zealand) carried only two haplotypes: Arc1 and Arc2 [35]. It also showed that Australian dingoes carried only haplotype A29 or haplotypes differing from A29 by a single mutation, indicating that the dingo population was founded from a small number of dogs carrying a single mtDNA haplotype (A29). Importantly, all three haplotypes are typical for East Asian dogs: Arc2 and A29 are absent and Arc1 rarely found west of the Himalayas [9]. Furthermore, two New Guinea singing dogs (NGSDs; a feral dog from the New Guinean highlands, close in morphology and behaviour to Australian dingoes but clearly distinguishable [36]) were shown to carry haplotypes A29 and A79 (which differs by one substitution from A29). Since A29 is also found among East Asian dogs, an origin from domestic dogs seems clear for these two wild populations.
...
This study shows a distinct pattern in the geographical distribution of the two Polynesian dog mtDNA haplotypes Arc1 and Arc2, and the dingo and NGSD founder mtDNA haplotype A29, with a total frequency of 12 per cent in Southern China, 17 per cent in southeast Asia and 50 per cent in Indonesia, but complete absence in samples from Taiwan and the Philippines. This gives a clear indication that Polynesian dogs as well as dingoes and NGSDs trace their ancestry back to South China through Mainland Southeast Asia and Indonesia. Thus, there is no indication that these dogs were introduced via Taiwan and the Philippines together with the expansion of the Neolithic culture and Austronesian languages, as suggested in some theories about Polynesian origins.
So it clearly indicates an origin in Southern China and a route via Southeast Asia and Indonesia. Also that they are not a seperate species. So for several years now genetic analyses do not support a classification as a seperate species albeit the term Canis hallstromi is still in use. Morphological data also doesn't support it, to a huge degree due to the high morphological range of the domestic dog and as even Matznick states the physical capabilities of the NGSD are not as unique as often stated, since the Lundehund is very similar in that regard. Furthermore I know of no source that ever stated that NGSD males have an actual breeding cycle and an annual breeding cycle for females is not unknown among the domestic dog (and NGSD females can have three heats per year), it is simply not the most common. Furthermore reduced relative brain size, hypertrophied tail form (curled tail), reduction of facial expression with a stronger emphasis on phonetic expression and a less sophisticated social system when compared to examined grey wolves are all characteristics of the domestic dog and they are shared by the NGSD. So neither genetic nor morphological data support a classification as a separate species from the dog. In addition a big part of the few founding dogs of the captive population already came from humans so simply classifying them as wild is also on shaky grounds, but this last argument we should not use.
Now the biological definition of species also doesn't fit, especially not when all of the wild population, or at least the major part, has mixed with other dogs. Two seperate species don't mix so readily and so extensively. I know that at least some authors state that all members of the genus Canis can produce fertile offspring. I don't know of the basis of that statement only that it was proven for lupus, aureus, simensis, rufus, latrans and possibly lycaon (if that is a seperate wolf-species), and the gray wolf and domestic dog are considered one species due to proven voluntary mating when sufficent number of partners where there (so neither a shortage of dogs or wolves), this was under captive conditions. And an average unrestricted fertility of the hybrids and no sudden increase in genetic disorders even when only bred among each other since the first generation of hybrids was only proven when you mix gray wolves with domestic dogs.
Now, as far as is known to me the taxonomic classification of the NGSD is according to Mammal Species of the World the NGSD is under Canis lupus dingo http://www.bucknell.edu/msw3/browse.asp?id=14000751 and the term hallstromi is listed among the synonyms which also contains many old names of the dingo. Also the entry for c.l. dingo also stated as a comment "domestic dog" which is also in the entry for c.l. familiaris http://www.bucknell.edu/msw3/browse.asp?id=14000752 . Under the entry for Canis lupus http://www.bucknell.edu/msw3/browse.asp?id=14000738 it is stated Includes the domestic dog as a subspecies, with the dingo provisionally separate--artificial variants created by domestication and selective breeding (Vilá et al., 1999; Wayne and Ostrander, 1999; Savolainen et al., 2002). Although this may stretch the subspecies concept, it retains the correct allocation of synonyms. Corbet and Hill (1992) suggested treating the domestic dog as a separate species in SE Asia. However the seperate species argument doesn't held up for the reasons I have mentioned.
So I am sure that the current classification of the NGSD is c.l. dingo and since the NGSD has the mtDNA type A29 like the Australian dingo that is as far as I know regarded as the current justification for its subspecies classifcation. However, it is clearly seperate from the Australian and Thai version of the dingo due to morphology and also phonetic and social characteristics (all based on the captive population of course), therefore it needs a seperate article in my eyes just like most breeds of dog have, also the topic has the necessary amount of material and holds enough interest for a seperate article.
Due to all this: A classification as a subspecies is basically just as arbitrary as that for race or breed and when we cannot get rid of the c.l. dingo in the taxobox it has to stay. But the article needs to state the other terms that were once and are often still in use along with the published material on genetic and morphological analysis. I hope I could help. --Inugami-bargho (talk) 19:47, 7 December 2011 (UTC)
Taxonomy of Canis in general
Well, the taxonomy of Canis is a nightmare, with species, semi-species and subspecies blending neatly into each other, and likely several paraphyletic groups at the species level. Just to top it off, the species barriers seem to be social (pre-zygotic) rather than genetic, and with domestication humans are having a hand in it as well. I think it would be useful to accnowledge that in Canis there are possibly no good species, they are as much human constructs as a biological reality. Petter Bøckman (talk) 15:45, 12 November 2011 (UTC)
I don't know exactly what you mean, but I've noticed a few things that cause problems for Wikipedia. First of all, even though the dog is a domesticated wolf, now Canis lupus isn't doesn't jive with the English (just to mention one language) "wolf" because dogs aren't wolves and dogs are Canis lupus too, so unless we merge the articles wolf and dog we have no article for Canis lupus in general anymore. Next, the dog is now two taxa even though one is the other gone feral or wild, Canis lupus familiaris and C.l.dingo, which causes a mess for wikipedia in terms of what to put where and how much each article should cover. Next, the Golden Jackal on the one hand and the other two jackals on the other turn out now to belong to two differnet branches, so the article Jackal has headaches. All existing red wolves, and maybe all Canis lupus rufus or Canis rufus turn out to be Coydogs, but the editors of that article can't handle the truth so it's a mess. The "Canis latrans" around here NYS are maybe 80% latrans, tops, it turns out, the rest is lupus, mostly wolf but partly dog. I could go on, but wikipedia is steadily sorting through it all, it's kinda fun to watch/be a part of, sort of a puzzle. But this NGSD article has been a headache from day one. Chrisrus (talk) 05:00, 13 November 2011 (UTC)
Oh, it is a puzzle indeed! There are at least three distinct problems mixed in together:
Paraphyly, how do you treat paraphyletic, but obviously useful groups (like "wolf")? Even worse, what do you do with polyphyletic (hybrid) species?
Language vs phylogeny, what do you do when the two don't match?
The point I was trying to make (and which I think apply to the NGSDs) is thet there is quite possibly no one correct solution here. Petter Bøckman (talk) 08:43, 13 November 2011 (UTC)
This is, of course, not specific to Canis, but is a general problem in groups where geographically and morpologically distinct populations are quite capable of inter-breeding and do so regularly. The number of species of Ophrys (bee orchids) recognized in reliable sources varies from about 10 to 375! There are two different issues in Wikipedia: which entities should have their own articles, which should surely be determined by interest and the amount of information available, and what (if anything) should be put in taxoboxes. I'm a little concerned that some editors (not those who have commented here) seem to want the latter to determine the former. Peter coxhead (talk) 10:01, 13 November 2011 (UTC)
A huge part of the problem is also the classification of the domestic dog and especially in the english speaking world it seems hard to accept for many that domestic dog and gray wolf are the same species. Partly this comes from the inconsistency on what is regard a typical characteristic of the domnestic dog (e.g. many see lack of regurgitating of food as normal while others as a sign of degeneration since other dogs still show it perfectly [even males]) as well as idological reasons (e.g. the arbitrary seperation of nature and culture). So yeah, we probably should have a different article for canis lupus than the current one, but I don't know whether such an article would ever make it. Does anybody here know whether such an article would find support?--Inugami-bargho (talk) 19:47, 7 December 2011 (UTC)
For some time, a search for Canis lupus linked to the article Wolf. Then, when it was pointed out that the article wasn't talking about the whole of Canis lupus per se when it said things such thing such as their being less adaptable to civilized times than coyotes and many other such statements, a hatnote was added to indicate that the article was about wolves, not dogs, which are also canis lupus. Then, for awhile, the hatnote was removed and Canis lupus searchers were sent directly to Subspecies of Canis lupus, I suppose because it was the closest thing to an article about Canis lupus in general and if such an article as you describe would be created, it might incorporate that one. Then, perhaps reasoning that "Canis lupus" searchers were probably looking for the Wolf article, Canis lupus searchers were direct back to the "wolf" article again, but hatnote was re-added and changed so that the if anyones clicks on the hatnote Canis lupus link at the top of the wolf article, s/he is sent to the subspecies article. That's how it stands today.
The dog article and every doginfobox was at first assigned C.l.familiaris and there was only one article, Dingo, which was also Canis lupus dingo. Then it was noticed that halstromiAustralian Dingo" and Canis lupus dingo was created to embark all dogs so classified. Meanwhile, the article "Dog" was changed so that it would cover both c.l.familiaris and c.l.dingo, and these two were tied together using the brackets around the term "[domestic dog]", in imitation of the way it's done in MSW3, but no one seems to understand" because editors regularly change it because they don't understand why the domestic dog and the dingo are both the "[domestic dog]" in the chart. So some other way to make MSW3's point in the chart seems needed. Chrisrus (talk) 01:17, 8 December 2011 (UTC)
What a mess! The problem here is that we seek to make articles to cover specific phenomena, and these phenomena aren't clear-cut phylogenetic units. As a zoologist I very much like to see articles follow scientific naming, but in this case case I'm tempted to say "screw phylogeny" and stick to the vernacular expressions. There really should be an article on wolf, on dog, on dingo, on domestic dog etc. The relevant phylogenetic information clearly should be given in each, but should not dictate what articles to write. (This is why I prefer to stick to wild animals, things get difficult when humans start meddling) Petter Bøckman (talk) 08:21, 8 December 2011 (UTC)
Oh, great. This is one of the reasons I don't get why the english wikipedia doesn't have a peer review system that prevents that every edit of a site can be seen immedidiately. Ok, lets see.
I think the main problem was the original wolf article because it wasn't in accordance with the facts. I just checked it and its worse than I imagined since it isn't even in accordance with the sources it states (e.g. it references MSW3 for the classifcation of subspecies but excludes familiaris and dingo nonetheless). It is the old problem of generalization of things that are not the same. So basically that one would need to be changed first but I doubt we can do it if there is such massive resistance to it. I think a new article should be done and reviewed before published. The chart implies that dingoes don't have the traits typical for domestic dogs. The main problem we would face, based on what you wrote, would be the opposition against stating that the domestic dog is a gray wolf. In addition the article on the domestic dog needs to be seriously reviewed. But that is such a huge project that we would need several dozen authors.--Inugami-bargho (talk) 08:25, 8 December 2011 (UTC)
As an outsider in "dog matters", it seems to me that Petter Bøckman is right. What readers want is articles on topics they expect to be different, i.e. wolf, dingo, domestic dog, etc. These terms have clear meanings regardless of the taxon referred to. By the way, I think it's wrong to say that "the domestic dog is a gray wolf". The domestic dog and the gray wolf may belong to the same species, but this doesn't make their ordinary language meanings the same. (A similar issue of language arises regularly at Ape and Primate, where people try to impose the usage "apes" = "hominoids including humans", instead of reporting this as one of the usages attested to in reliable sources.)
I think that we also sometimes run into trouble with plant articles by not separating out cultivated and wild forms. Popular ornamental plants which have many cultivars and have probably been been massively crossed, for example, don't fit well into taxonomic hierarchies. Peter coxhead (talk) 11:16, 8 December 2011 (UTC)
No offense but "ordinary language meanings" doesn't count. Wikipedia is basically a lexicon and not a forum where people get their expectations confirmed. Furthermore you are wrong in thinking that the canine terms you mentioned have clear meanings. I know of a t least three definitions for dingo as well as three different usages regarding what dogs are dingoes and what not. The same with the Australian term of "wild dog". It is the same with the term domestic dog, for some this includes all canids with a history of domestication so including the dingo, for others the term only means pet varietes and either including or excluding working dogs, for others again feral dogs are domestic dogs but for others not, others call feral dogs stray dogs or wild dogs, it is the same with horses. To go deeper, if we would take such "ordinary language meanings" into account we would have to classify British Bulldogs as perfectly healthy because owners and breeders might be offended, despite the fact that from an objective point of view they are cripples. So we cannot afford to take "ordinary language meaning" into account when we want to have good and reliable articles.--Inugami-bargho (talk) 07:11, 10 December 2011 (UTC)
The idea of having one article about Canis lupus the taxon is maybe a good one. It be Subspecies of Canis lupus and about the history of the term, how it used to be just wolves and then started to cover dogs when it was determined that they are domesticated from wolves, and so on. The idea that we should merge the articles Wolf and dog just because taxonomy has done so just not reasonable, neither in English nor German nor any of the other languages. Chrisrus (talk) 13:53, 10 December 2011 (UTC)
I think your may have misunderstood my stance, Inugami-bargho. Of course we need an article on Canis lupus in it′s full meaning. It just does not need to be the article "Wolf". Wikipedia has plenty of room, there′s room for both, and with links and room to explain the changing meaning of terms, no reader needs to feel confused over where to find what on Wikipedia. Petter Bøckman (talk) 19:22, 10 December 2011 (UTC)
And I think you misunderstand me. When you say wolf in science you mean Canis lupus unless you first stated what subspecies you are talking about. For the respective subspecies you need to name the non-taxonomic name. That in english speaking countries they often exclude dogs from that is because there is such a problem for most people in recognizing that both are the same species, as far as I know that is due to the phenomenon that most dog researchers have no idea about wolves and most wolf-researchers do not even own a dog. So yes the article "wolf" would have to be about the whole of Canis lupus especially since the current article implies that all these wild populations are the same, but there are differences among them as well (e.g. North American wolves were often eradicated after only 20 years of European colonization while in Europe it was only successfull in the 19th century and even then only slowly). Therefore we need this one article stating the basic facts and including the current accepted list of subspecies and from there links to articles on each subspecies, including articles (if possible) on domestication, mythology and the like. This article can actually be very short since more detailed information on the various wolf-populations can be covered in the various subspecies entries. I don't see how that could be confusing. --Inugami-bargho (talk) 08:39, 13 December 2011 (UTC)
Your suggestion would have been the sensible one if phylogeny was the main criterium for organizing articles. However, phylogeny is but one of several criteria, and the size of any article depends on what people want to put in it. A lot of people are interested in wolfs, no matter how short the article could have been after your suggestion, it would soon bloat out and become the de facto main article. Your suggestion that wolfs really are not a single species is interesting, but would violate WP:OR and WP:WEIGHT until the species is formally split, and I don't see that happening anytime soon. If we do not insist on fighting a never ending uphill battle, the sensible thing to do is to have an article on Wolf (Canis lupus) with all that normally goes into a species article, and treat the subspecies and the cultivar (dog) is small sections (a couple of paragraphs) with links to their respective main articles. Petter Bøckman (talk) 10:42, 13 December 2011 (UTC)
It doesn't matter whether people would bloat the article or not. You can't just give in to pressure because of what people want, when you go by that criteria the wolf article would contain info about the thylacine or the maned wolf as well. And I didn't say that wolves are not a single species. There is only one species named gray wolf and it consists of several variations that differ from each other, e.g. polar wolves are on average much bigger and heavier than Italian wolves but they are still the same species. " If we do not insist on fighting a never ending uphill battle, the sensible thing to do is to have an article on Wolf (Canis lupus) with all that normally goes into a species article, and treat the subspecies and the cultivar (dog) is small sections (a couple of paragraphs) with links to their respective main articles." That was what I was saying the whole time. However we would need to have the actual full range meaning in the size it would be from e.g. the weight of a toy dog to that of a mastino and there we could have a problem. We would need a fool proof argument for the inclusion of the domestic dog since many will probably oppose that no matter what behavioral and genetic examinations tell us. Anyway the best approach would be to first create an appropriate article on the user page of an author, when that one is sufficient it can be published and we should request for a protection of the article. Maybe even have it locked.--Inugami-bargho (talk) 14:36, 16 December 2011 (UTC)
Are you talking at cross purposes? There should be one article for Dog, one for Wolf, and one for Dingo, so the criticism about "giving in to what people want" does not apply unless you are arguing that all of these be merged into one Canis lupus article, which alternately seem to be advocating and then not advocating. But if you really want to merge these into one Canis lupus article, please try that on German Wikipedia first and see what happens there. I can't believe that's what you are really suggesting, but that's what the Peters and I were arguing against, and what you seemed to be arguing for when you object to "ordinary language meanings" and "giving people what they want". If on the other hand you'd like to have an additional article about Canis lupus the taxon, please go ahead and create it out of the existing article Subspecies of Canis lupus, as that's the existing article about that referent and could be expanded, for example it presently lacks a history of the taxon, starting with Linnaeus, and other things. Then you could move that article to Canis lupus and let the wolf and dog articles continue to just talk about wolves and dogs as that's quite enough for each of them. Chrisrus (talk) 18:35, 16 December 2011 (UTC)
Please everyone please read the article Red Wolf and its discussion page TALK:Red wolf and comment here and/or there. There are some similar issues and some different ones and room for improvement and lessons to be learned for the wider project. Chrisrus (talk) 18:05, 11 December 2011 (UTC)
First, there are some debates, not all of which are closed, on the talk page that may need resolution.
But to my mind, the problem is this: about half-way through the Taxonomy section the reader is told that all existant red wolves are coywolves, based on DNA evidence. The lead says nothing about this, calling it simply a "canid", which isn't particularly enlightening. I can't evaluate the genetic evidence, but it seems that for at least a decade the evidence has been conclusive that two branches on the tree of life in this case have merged into one, and it seems to me that if that's true and the last word has been said about that, it should be in the lead because it goes to the heart of what this animal fundementally is, which is a rare and to my mind significant thing, because even though Canis hybrids and others aren't so rare, they rarely form "true-breeding" sustained populations, and in no other case has mammology ever recognized one as a taxon. So it seems to me one of the most important things about it, what is fundementally is, is not in the lead, and that the way the Taxonomy section is done undue weight seems to lie on the theory that it's not a coywolf if that's no longer a tenable theory. But as I'm not qualified to evaluate the various points of view on the subject I wanted to ask before changing the lead based on only the second half of the Taxonomy section. Chrisrus (talk) 00:36, 12 December 2011 (UTC)
First off, there's no one good solution to the species problem of the red wolf. No form of scientific classification really handle ambiguity well. We will have to accept that we are left with a "best of a bad situation" as the optimum solution. As long as everyone realize that whatever solution is found, it will have flaws, then we should be able to find consensus.
From a phylogenetic POW, the problem with the red wolf is not really that it appears to be a good species ecologically, but that wolf appears to be one too. Phylogenetic nomenclature is based on species neatly splitting in two and the ancestor species then going extinct. This assumption if of course just a model, not real life, and every now and then we run into situations where this assumption very clearly do not hold. There are quite a few instances of paraphyletic species out there, and hybridization of recently diverged species may be more common in mammals than assumed, our own speciation from the chimpanzee being a point in question. We just need to find how to best handle this situation in the taxobox, and make sure the whole mess is explained clearly in the text.
As far as I can see, the red wolf as a hybrid species seems fairly conclusive. Then again, hybrid species are species too, or can at least give rise to species if allowed to evolve. At what point do a species cease to be "just a hybrid" and start becoming a species of it's own? Petter Bøckman (talk) 18:32, 12 December 2011 (UTC)
Thanks for all you've done with regard to this. Please, don't stop there. As I said before (scroll up), the problem is the "lede" sentence(s). We need to state in reader-friendly language what it is. How about "a hybrid "species"" or a descriptive phrase to that effect? "Canid" just isn't very enlightening. Don't worry so much about taxonomy. Chrisrus (talk) 23:31, 6 February 2012 (UTC)
The terminology would be "suspected/presumed/hypothesized/whatever hybridogenic species" (or hybridogenicspecies if you prefer it blue). Dysmorodrepanis (talk) 23:33, 15 June 2012 (UTC)
Ok, then, please do go there and make that change because not just the body but also the lead should say in as upfront and clear a way as appropriate that this is a hybrid species or a hybridogenic species. Because it still takes to much digging for the reader to understand this. Chrisrus (talk) 00:00, 16 June 2012 (UTC)
Please if you would do have a look at Talk:Coywolf re: Redwolf, and comment as you would. I will add an opposite message there. Chrisrus (talk) 17:22, 6 September 2012 (UTC)
Blackbacked and Side-striped Jackals:
Dear WikiProject Tree of Life page watcher;
I found this uncited section here Canid_hybrid#Genetic_considerations. We could, I suppose, delete it on the grounds that i's uncited, but the context kinda needs it and it sounds like it was written by someone who knows what they are talking about and seems logical. So instead, I'm trying to cite it. But I'm not really able without some help from an expert. It needs checking and citing or correcting if the numbers something's wrong. Could this here: http://www.ncbi.nlm.nih.gov/pmc/articles/PMC53565/pdf/pnas01030-0152.pdf be used to cite the part about the Black-backed Jackal and the Side-striped Jackal?
Thank you for your kind attention! Chrisrus (talk) 06:10, 21 December 2011 (UTC)
I've found some sources for the chromosome numbers, but not yet for hybridization. Petter Bøckman (talk) 08:46, 21 December 2011 (UTC)
Please, expert needed to explain MSW3's statements on the taxon Canis lupus dingo page has only two words in the comments section, placed in brackets, here: http://www.bucknell.edu/MSW3/browse.asp?id=14000751. This is also true of their Canis lupus familiaris page. What does this mean? Please, any/every expert, what does this mean?
Of course, before you answer, you should read the exlanation that there seems to be on their Canis lupus page, here: http://www.bucknell.edu/MSW3/browse.asp?id=14000738. It says many things about the taxonomic questions about this species, but it says this about this matter:
(Canis lupus) includes the domestic dog as a subspecies, with the dingo provisionally separate--artificial variants created by domestication and selective breeding (Vilá et al., 1999; Wayne and Ostrander, 1999; Savolainen et al., 2002). Although this may stretch the subspecies concept, it retains the correct allocation of synonyms. Corbet and Hill (1992) suggested treating the domestic dog as a separate species in SE Asia. Synonyms allocated according to Ellerman and Morrison-Scott (1951), Mech (1974), and Hall (1981).
Maybe you should also have a look at the article Canis lupus dingo, which is not the same as the Australian Dingo article, and not overequate the Australian native dog with all the dogs covered by the taxon Canis lupus dingo.
Thank you for your kind attention to this matter. Chrisrus (talk) 04:43, 16 February 2012 (UTC)