.mw-parser-output .hidden-begin{box-sizing:border-box;width:100%;padding:5px;border:none;font-size:95%}.mw-parser-output .hidden-title{font-weight:bold;line-height:1.6;text-align:left}.mw-parser-output .hidden-content{text-align:left}You can help expand this article with text translated from the corresponding article in Polish. (June 2015) Click [show] for important translation instructions.
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He studied at the University of Jena under Ernst Haeckel, obtaining his doctorate for studies of embryo cleavage in Hydra. His later work Hydra - Eine anatomisch-entwicklungsgeschichtliche untersuchung, in English "An anatomical-evolutionary investigation of Hydra" (Leipzig, Wilhelm Engelmann, 1872) is a classic, still quoted monograph which has implications for evolution theory.
Die Entstehung des Annelids aus der Larve von Lopadorhynchus. Nebst bemerkungen über die Entwicklung anderer Polychaeten, 1886 - On the formation of annelids from the larvae of Lopadorhynchus.[2]