| Ademosynidae Temporal range:
| |
|---|---|
Scientific classification 
| |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Coleoptera |
| Suborder: | incertae sedis |
| Family: | †Ademosynidae Ponomarenko, 1968 |
Ademosynidae is an extinct family of beetles, known from the Late Triassic to Early Cretaceous. There are at least six genera and 11 species in Ademosynidae.[1][2][3][4] Members of the family were small oval beetles, with a length generally smaller than 1 cm. Characteristics of the family include a pronotum without anterior angles and a rounded anterior margin, and elytra with 9–12 punctate striae. The systematic position of the family within the order Coleoptera is currently uncertain: they can be considered early examples of the suborder Polyphaga, or unusual examples of Archostemata.[3]
Genera
[edit]The family Ademosynidae is restricted to the following six genera and 11 species in Yan, Beutel and Ponomarenko (2017):[3]
- † Ademosyne Handlirsch, 1906
- †Ademosyne bacca Ponomarenko, 1969 – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
- †Ademosyne elliptica Ponomarenko, 1969 – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
- †Ademosyne kirghizica Ponomarenko, 1969 – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
- †Ademosyne major Handlirsch, 1906 – Blackstone Formation, Australia, Late Triassic (Norian)
- † Cephalosyne Ponomarenko, 1969 – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
- †Cephalosyne capitata Ponomarenko, 1969
- † Dolichosyne Ponomarenko, 1969 – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
- †Dolichosyne confragosa Ponomarenko, 1969
- †Dolichosyne rostrata Ponomarenko, 1969
- †Dolichosyne sulcata Ponomarenko, 1969
- † Gnathosyne Ponomarenko, 1969 – Tologoi Formation, Kazakhstan, Late Triassic (Norian)
- †Gnathosyne akkolkensis Ponomarenko, 1969
- † Petrosyne Ponomarenko, 1969 – Kyrgyzstan, Early Jurassic (Pliensbachian)
- †Petrosyne liassica Ponomarenko, 1969
- † Sphaerosyne Ponomarenko, 1969 – Zaza Formation, Russia, Early Cretaceous (Aptian)
- †Sphaerosyne globosa Ponomarenko, 1969
The following genera and species are also included in the family according to Fossilworks, but are excluded or not mentioned in Yan, Beutel and Ponomarenko (2017):[2]
- † Ademosynoides Dunstan, 1923
- †Ademosynoides abnormis Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †Ademosynoides alternata Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †Ademosynoides angusta (Tillyard, 1916) – Blackstone Formation, Australia, Late Triassic (Norian)
- †Ademosynoides antarctica Zeuner, 1959 – Mount Flora Formation, Antarctica, Middle Jurassic (Bathonian)
- †Ademosynoides japonicus Fujiyama, 1973 – Mononoki Formation, Japan, Late Triassic (Carnian)
- †Ademosynoides magnifica Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †Ademosynoides minor (Handlirsch, 1906) – Blackstone Formation, Australia, Late Triassic (Norian)
- †Ademosynoides obtusa (Tillyard, 1916) – Blackstone Formation, Australia, Late Triassic (Norian)
- †Ademosynoides striatella Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- † Atalosyne Ren, 1995 – Lushangfen Formation, China, Early Cretaceous (Aptian)
- †Atalosyne sinuolata Ren, 1995
- † Chaocoleus Ponomarenko, Yan & Huang, 2014 – Yinping Formation, China, Permian (Capitanian)
- †Chaocoleus limnebius Ponomarenko, Yan & Huang, 2014
- † Eremisyne Wang, 1998 – Xiazhuang Formation, China, Early Cretaceous (Aptian/Albian)
- †Eremisyne xiazhuangensis Wang, 1998
- † Grammositus Dunstan, 1923 (also spelled Grammositum) – Blackstone Formation, Australia, Late Triassic (Norian)
- †Grammositus bilineatus Dunstan, 1923
- †"Ademosyne" adunca Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" arcucciae Martins-Neto and Gallego, 2006 – Los Rastros Formation, Argentina, Late Triassic (Carnian)
- †"Ademosyne" australiensis Tillyard, 1916 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" brevis Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" cameroni Tillyard, 1916 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" congener Tillyard, 1916 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" curvata Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" elongatus Martins-Neto and Gallego, 2006 – Los Rastros Formation, Argentina, Late Triassic (Carnian)
- †"Ademosyne" hexacostata Martins-Neto and Gallego, 2006 – Los Rastros Formation, Argentina, Late Triassic (Carnian)
- †"Ademosyne" intermedia Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" lata Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" llantenesensis Brauckmann et al., 2010 – Llantenes Formation, Argentina, Late Triassic (Norian)
- †"Ademosyne" olliffi (Handlirsch, 1906) – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" parva Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" polyzetete Ponomarenko, 2011 – Lunz Formation, Austria, Late Triassic (Carnian)
- †"Ademosyne" prisca Riek, 1974 – Molteno Formation, South Africa, Late Triassic (Carnian)
- †"Ademosyne" punctata Tillyard, 1916 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" punctuada Martins-Neto and Gallego, 2006 – Los Rastros Formation, Argentina, Late Triassic (Carnian)
- †"Ademosyne" ramocostata Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" reducta Riek, 1976 – Molteno Formation, South Africa, Late Triassic (Carnian)
- †"Ademosyne" rosenfeldi Brauckmann et al., 2010 – Llantenes Formation, Argentina, Late Triassic (Norian)
- †"Ademosyne" rugulosa Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" sibirica Rohdendorf, 1961 – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian (Wuchiapingian/Changhsingian)
- †"Ademosyne" speciosa Riek, 1974 – Molteno Formation, South Africa, Late Triassic (Carnian)
- †"Ademosyne" umutu Martins-Neto and Gallego, 2009 – Los Rastros Formation, Argentina, Late Triassic (Carnian)
- †"Ademosyne" vittamargina Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- †"Ademosyne" wianamattensis Tillyard, 1918 – Ashfield Shale, Australia, Middle Triassic (Pelsonian/Illyrian)
References
[edit]- ^ "Ademosynidae". GBIF. Retrieved 2019-10-21.
- ^ a b "The Paleobiology Database, family Ademosynidae". Retrieved 2019-10-21.
- ^ a b c Yan, Evgeny V.; Beutel, Rolf G.; Ponomarenko, Alexander G. (2017). "Ademosynidae (Insecta: Coleoptera): A new concept for a coleopteran key taxon and its phylogenetic affinities to the extant suborders". Palaeontologia Electronica. 20. doi:10.26879/739.
- ^ Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. Bibcode:2011ZooK...88....1B. doi:10.3897/zookeys.88.807. PMC 3088472. PMID 21594053.