Family of gastropods
Bulimulidae is a taxonomic family of medium-sized to large, air-breathing, tropical and sub-tropical land snails , terrestrial pulmonate gastropod molluscs in the superfamily Orthalicoidea .[ 2]
Distribution of species in the family Bulimulidae includes Ecuador (9 genera)[ 3] and other South American countries. Some species also occur in North America.
The family's oldest fossil record dates from the late Cretaceous of Brazil (Itaboraí Basin).[ 4] [ 5]
Members of this family have a haploid chromosome number between 26 and 30 (according to the values in this table).[ 6]
Shells of species within Bostrycinae have a smooth protoconch and the genital organs feature a relatively long penis sheath (ca. 1/4–1/6 total phallus length).[ 7]
Previously, the members of the Orthalicidae were also included in this family, as the subfamily Orthalicinae, and the taxa listed here were placed in their own subfamily, the Bulimulinae.[ 8]
Bulimulinae was placed in the family Orthalicidae according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005) .[ 9]
The subfamily Bulimulinae included 3 tribes:[ 9]
Tribe Bulimulini Tryon, 1867 - synonyms: Bulimidae Guilding, 1828 (inv.); Berendtiinae P. Fischer & Crosse, 1872; Bothiembryontidae Iredale, 1937
Tribe Odontostomini Pilsbry & Vanatta, 1898 - synonym: Tomogeridae Jousseaume, 1877
Tribe Simpulopsini Schileyko, 1999 Breure et al. (2010)[ 1] moved Bothriembryon to Placostylidae (since 2012 as Bothriembryontidae), elevated Bulimulinae to Bulimulidae and Odontostomini to Odontostomidae .[ 1]
Breure & Romero (2012)[ 10] confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae .[ 10] There are three subfamilies within Bulimulidae:[ 10]
subfamily Bulimulinae Tryon, 1867
subfamily Peltellinae Gray, 1855
subfamily Bostrycinae Breure, 2012[ 10] [ 7] Genera in the family Bulimulidae include:
subfamily Bulimulinae Tryon, 1887
(synonym: Berendtiinae P. Fischer & Crosse, 1872 ) Anctus E. von Martens, 1860
Auris Spix, 1827
Berendtia Crosse & P. Fischer, 1869
Bocourtia Rochebrune, 1882
Bulimulus Leach, 1814 - type genus of the subfamily Bulimulinae[ 9] [ 3] [ 10]
Cochlorina Jan, 1830
Graptostracus Pilsbry, 1939
Kora Simone, 2012[ 11]
Lopesianus Weyrauch, 1958
Naesiotus Albers, 1850[ 12] [ 13] [ 3] [ 10]
Neopetraeus E. von Martens, 1885
Newboldius Pilsbry, 1932
Otostomus H. Beck, 1837
Oxychona Mörch, 1852
Protoglyptus Pilsbry, 1897[ 14]
Pseudoxychona Pilsbry, 1930
Rabdotus Albers, 1850[ 10]
Scutalus Albers, 1850[ 15] [ 3] [ 10]
Spartocentrum Dall, 1895
Sphaeroconcha Breure, 1978
Stenostylus Pilsbry, 1898[ 13] [ 3] [ 14]
Suniellus Breure, 1978[ 14] subfamily Peltellinae subfamily Bostrycinae Unassigned within Bulimulidae Bulimus Bruguière, 1789 (temporary name)
† Itaborahia Maury, 1935[ 4]
† Oreoconus D. W. Taylor in McKenna et al., 1962
† Palaeobulimulus Parodiz, 1949
Stapafurdius Simone, 2021
† Tocobaga Auffenberg, Slapcinsky & Portell, 2015 Adzharia P. Hesse, 1933 : synonym of Bulimulus Leach, 1814 (junior synonym)
Antidrymaeus Germain, 1907 : synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
Atahualpa Strebel, 1910 : synonym of Thaumastus E. von Martens, 1860
Ataxellus Dall, 1912 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented asBostryx Troschel, 1847
Ataxus Albers, 1850 : synonym of Bostryx Troschel, 1847
Bilamelliferus Weyrauch, 1958 : synonym of Bostryx Troschel, 1847 (junior synonymy)
Cochlogena Férussac, 1821 : synonym of Bulimulus Leach, 1814 (junior synonym)
Diaphanomormus Weyrauch, 1964 : synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
Elatibostryx Weyrauch, 1958 : synonym of Bostryx Troschel, 1847
Floreziellus Weyrauch, 1967 : synonym of Bostryx Troschel, 1847 (junior synonymy)
Geoceras Pilsbry, 1896 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
Globulinus Crosse & P. Fischer, 1875 : synonym of Rabdotus Albers, 1850
Goniognathmus Crosse & P. Fischer, 1875 : synonym of Drymaeus Albers, 1850 (junior synonym)
Hamadryas Albers, 1850 : synonym of Drymaeus Albers, 1850
Hannarabdotus Emerson & Jacobson, 1964 : synonym of Rabdotus Albers, 1850
Kionoptyx Haas, 1966 : synonym of Bostryx Troschel, 1847 (junior synonymy)
Kuschelenia Hylton Scott, 1951 : synonym of Bocourtia (Kuschelenia) Hylton Scott, 1951 represented as Bocourtia Rochebrune, 1882
Leptodrymaeus Pilsbry, 1946 : synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
Leptomerus Albers, 1850 : synonym of Bulimulus Leach, 1814
Leptomormus Weyrauch, 1958 : synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
Liostracus [sic]: synonym of Leiostracus Albers, 1850 (misspelling of original genus, Leiostracus Albers, 1850 )
Loboa Ihering, 1917 : synonym of Bulimulus Leach, 1814
Mormus E. von Martens, 1860 : synonym of Drymaeus Albers, 1850
Multifasciatus Weyrauch, 1958 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
Naesiotellus Weyrauch, 1967 : synonym of Bostryx Troschel, 1847 (junior synonym)
Navicula Spix, 1827 : synonym of Cochlorina Jan, 1830
Nesiotes E. von Martens, 1860 : synonym of Naesiotus Albers, 1850
Obstrussus Parodiz, 1946 : synonym of Naesiotus Albers, 1850
Olinodia Dall, 1920 : synonym of Naesiotus (Reclasta) Dall, 1920 represented as Naesiotus Albers, 1850
Orphaicus Schaufuss, 1869 : synonym of Thaumastus E. von Martens, 1860
Orphnus Albers, 185 0: synonym of Thaumastus E. von Martens, 1860
Orthotomium Crosse & P. Fischer, 1875 : synonym of Rabdotus Albers, 1850
Pachyotus H. Beck, 1837 : synonym of Auris Spix, 1827
Pachytholus Strebel, 1909 : synonym of Thaumastus E. von Martens, 1860
Pampasinus Weyrauch, 1958 : synonym of Bostryx (Platybostryx) Pilsbry, 1896 : synonym ofBostryx Troschel, 1847
Peronaeus Albers, 1850 : synonym of Bostryx Troschel, 1847
Phenacotaxus Dall, 1912 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
Plecochilus Agassiz, 1846 : synonym of Plekocheilus Guilding, 1827 (unjustified emendation)
Pseudoperonaeus Weyrauch, 1958 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
Pseudorhodea Dall, 1895 : synonym of Rabdotus (Plicolumna) J.G. Cooper, 1895 represented as Rabdotus Albers, 1850
Pyrgus Albers, 1850 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
Rhabdotus [sic]: synonym of Rabdotus Albers, 1850 (incorrect subsequent spelling)
Rhaphiellus L. Pfeiffer, 1856 : synonym of Naesiotus Albers, 1850
Rimatula Parodiz, 1946 : synonym of Protoglyptus Pilsbry, 1897
Saeronia Dall, 1920 : synonym of Naesiotus Albers, 1850
Semiclausaria L. Pfeiffer, 1856 : synonym of Drymaeus Albers, 1850 (junior synonym)
Siphalomphix Rafinesque, 1833 : synonym of Bulimulus Leach, 1814
Sonorina Pilsbry, 1896 : synonym of Rabdotus (Leptobyrsus) Crosse & P. Fischer, 1875 represented as Rabdotus Albers, 1850
Spiroscutalus Pilsbry, 1932 : synonym of Scutalus Albers, 1850
Stenostoma Spix, 1827 : synonym of Anctus E. von Martens, 1860
Tatutor Jousseaume, 1887 : synonym of Thaumastus E. von Martens, 1860
Teneritia J. Mabille, 1898 : synonym of Spartocentrum Dall, 1895
Tholus Strebel, 1909 : synonym of Thaumastus E. von Martens, 1860
Xenothauma Fulton 1896:[ 17] [ 18] synonym of Scutalus Albers, 1850
Zaplagius Pilsbry, 1896 : synonym of Cochlorina Jan, 1830 This article incorporates CC-BY-3.0 text from the reference[ 7]
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