Brotia costula
Shell of Brotia costula (syntype at MNHN, Paris)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Superfamily:	Cerithioidea
Family:	Pachychilidae
Genus:	Brotia
Species:	B. costula
Binomial name
Brotia costula (Rafinesque, 1833)
Synonyms
see list

Brotia costula is a species of gastropod belonging to the family Pachychilidae.^{[1] [2]}

This species is found in Southeastern Asia.^[1]

Synonyms

• Brotia (Brotia) costula (Rafinesque, 1833)
• Brotia (Brotia) costula costula (Rafinesque, 1833)
• Brotia (Brotia) costula varicosa (Troschel, 1837)
• Brotia variabilis (Benson, 1836) (based on an unavailable name)
• Melania (Melanoides) soloensis K. Martin, 1905 † (junior subjective synonym)
• Melania (Melanoides) variabilis Benson, 1836 (superseded combination)
• Melania (Melanoides) variabilis var. microstoma G. Nevill, 1885 (original combination)
• Melania (Melanoides) variabilis var. pseudospinosa G. Nevill, 1885 (superseded combination)
• Melania (Melanoides) variabilis var. semilaevigata G. Nevill, 1885 (original combination)
• Melania (Melanoides) variabilis var. subspinosa G. Nevill, 1885 (original combination)
• Melania (Melanoides) variabilis var. subtuberculata G. Nevill, 1885 (original combination)
• Melania (Melanoides) variabilis var. subvaricosa G. Nevill, 1885 (original combination)
• Melania carolinae Gray in Griffith & Pidgeon, 1833 (junior synonym)
• Melania costula Rafinesque, 1833 (original combination)
• Melania hainesiana I. Lea, 1857
• Melania indica Souleyet, 1852 (junior synonym)
• Melania menkeana "Brot, 1875" (incorrect subsequent spelling of Melania menkiana; not qualifying as an emendation)
• Melania menkeana G. Nevill, 1885 (unjustified emendation of Melania menkiana I. Lea, 1843)
• Melania menkiana I. Lea, 1843 (junior synonym)
• Melania meukiana I. Lea, 1843 (incorrect original spelling; Melania menkiana is an emendation)
• Melania plicata I. Lea, 1838 (invalid: junior homonym of Melania plicata Menke, 1828; M. menkiana is a replacement name)
• Melania spinosa Hanley, 1854 (junior synonym)
• Melania variabilis Benson, 1836 (invalid: junior homonym of Melania variabilis Defrance, 1823)
• Melania variabilis var. aspera Hanley & Theobald, 1874 (invalid: junior homonym of Melania aspera Lesson, 1831)
• Melania variabilis var. cincta Hanley & Theobald, 1874 (invalid: junior homonym of Melania cincta I. Lea & H. C. Lea, 1851)
• Melania variabilis var. microstoma G. Nevill, 1885 (invalid: junior homonym of Melania microstoma I. Lea & H. C. Lea, 1851)
• Melania variabilis var. pseudospinosa G. Nevill, 1885 (junior synonym)
• Melania variabilis var. semilaevigata G. Nevill, 1885 (junior synonym)
• Melania variabilis var. subspinosa G. Nevill, 1885 (junior synonym)
• Melania variabilis var. subtuberculata G. Nevill, 1885 (junior synonym)
• Melania variabilis var. subvaricosa G. Nevill, 1885 (junior synonym)
• Melania varicosa Troschel, 1837